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few-leaf thistle, mountain thistle, Pacific fringe thistle, remote-leaf thistle, weak thistle

cobwebby thistle, snowy thistle, Venus thistle, western thistle

Habit Perennials, 20–150 cm, monocarpic; taprooted or polycarpic, perennating by runner roots. Biennials, 5–400 cm; taproots.
Stems

usually 1, erect, finely arachnoid-tomentose, sometimes villous with septate trichomes below nodes;

branches 0–10+, slender, usually arising in distal 1/2, ascending.

usually 1, thinly to densely gray- or white-tomentose, sometimes ± glabrate;

branches few–many, usually from above mid or near base in compact, moundlike dwarf plants, ascending to spreading.

Leaves

blades linear-oblong to oblanceolate or elliptic, 7–30 × 1–15 cm, unlobed and spinulose to dentate or shallowly to deeply pinnatifid, lobes well separated, linear to triangular-ovate, dentate to deeply lobed, main spines 2–5 mm, slender, abaxial faces green to gray, thinly to densely arachnoid-tomentose, sometimes glabrate, sometimes villous with septate trichomes along veins, adaxial green, glabrous;

basal sometimes present at flowering, sessile or winged-petiolate;

principal cauline mostly in proximal 1/2, winged-petiolate or sessile, bases narrowed, sometimes auriculate;

distal well separated, progressively reduced, becoming bractlike, often unlobed or less deeply divided than the proximal, sometimes spinier than proximal, bases often distally expanded and auriculate-clasping.

blades oblong–elliptic to oblanceolate, 6–40 × 1.5–10+ cm, shallowly to deeply pinnatifid, lobes usually rigidly spreading, undivided or with 1–2 pairs of coarse teeth or lobes, main spines 5–15 mm, both faces gray- to white-tomentose, sometimes ± glabrate or adaxial faces green, thinly arachnoid-tomentose;

basal sometimes present at flowering, petiolate or sessile and bases tapered, spiny-winged;

principal cauline much reduced distally, sessile, bases decurrent or not, as spiny wings;

distal much reduced, linear, ± bractlike.

Peduncles

(0–)2–15 cm.

1–30 cm.

Involucres

ovoid to hemispheric or campanulate, 1.5–2.5 × 1.5–3.5 cm, glabrous to arachnoid-floccose.

ovoid to spheric, 1.5–5 × 1.5–8 cm, arachnoid to ± loosely tomentose, often adjacent phyllaries connected by conspicuous arachnoid trichomes, sometimes glabrous or glabrate.

Corollas

creamy white to purple, 18–28 mm, tubes 7–12 mm, throats 5–12 mm, lobes 3.5–7 mm, style tips 4–6 mm.

white to lavender, pink, rose-purple, or red, 18–40 mm, tubes 8–18 mm, throats 5–7 mm, lobes 5–10 mm;

style tips 4–5 mm.

Phyllaries

in 6–8 series, subequal to strongly imbricate, green, linear to obovate (outer) to linear (inner), abaxial faces with inconspicuous glutinous ridge;

outer and middle bases appressed, margins entire to spinulose-dentate or broad, scarious, lacerate-dentate, spines absent or ascending to spreading, 1–2 mm;

apices of inner sometimes flexuous or reflexed, narrow, flat, entire or expanded, scarious, and lacerate-dentate.

in 7–10 series, subequal to strongly imbricate, green or stramineous to purple-tinged, linear to narrowly lanceolate, abaxial faces without glutinous ridge;

outer and mid bodies appressed, entire, apices deflexed to spreading or ascending, short-triangular to elongate, linear-acicular, spines spreading to reflexed, 1–10+ mm;

apices of inner erect, often flexuous, flat.

Heads

few–many, borne singly or in openly branched in corymbiform, racemiform, or paniculiform arrays on main stem and branches, sometimes also in distal axils, not closely subtended by clustered leaf bracts.

1–many in loose to tight clusters (barely raised above rosette in dwarf plants).

Cypselae

tan to dark brown, 4.5–5.5 mm, apical collars differentiated or not;

pappi 13–23 mm.

± brown, 5–6 mm, apical collars not differentiated;

pappi 15–30 mm.

2n

= 32.

Cirsium remotifolium

Cirsium occidentale

Distribution
from FNA
CA; OR; WA
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

Cirsium remotifolium occurs from the Coast Ranges and valleys of the Pacific Northwest to the western slopes of the Cascade and Klamath ranges, south in the California North Coast Ranges to the San Francisco Bay region. It is closely related to the C. clavatum complex of the Rocky Mountains region. Both have a similar growth habit and some forms variably express the character of broadly scarious, lacerate-toothed phyllary margins. Gray, in naming Cnicus carlinoides var. americanus, included as syntypes both California and Colorado specimens. F. Petrak (1917) treated both the West Coast plants and those of the Rocky Mountains as Cirsium subsect. Americana, recognizing C. remotifolium with several infraspecific taxa plus two other species, C. callilepis and C. amblylepis from the West Coast, and four additional species from the Rocky Mountains. A. Cronquist (1955) rejected Petrak’s subspecies, treating C. remotifolium in a restricted sense, limited to plants of Washington and Oregon without dilated phyllary tips, and circumscribed C. centaureae broadly to include the Rocky Mountains and West Coast plants with dilated phyllary tips. Because of the frequent presence of dilated phyllary tips in C. remotifolium in the restricted sense, Cronquist acknowledged the likelihood of past introgression with C. centaureae in the broad sense.

J. T. Howell (1960b) recognized three species: Cirsium remotifolium, C. acanthodontum, and C. callilepis, the latter with four varieties collectively corresponding to the West Coast representatives of C. centaureae (in the sense of Cronquist). Because of the great similarity of the various West Coast plants and their intergradation, I see no value in recognizing two or more species.

The West Coast and Rocky Mountains plants are clearly related, but are separated by the Great Basin region and there is little chance of current genetic interchange. As is often the case with American Cirsium, genetic enrichment from past hybridization with other nearby species within their respective areas has likely been fertile ground for evolutionary diversification. Different species have contributed genes in the Pacific states and in the Rockies. I have chosen to recognize two geographically-based species complexes, each with intergrading races here treated as varieties. I treat the West Coast plants as C. remotifolium and the Rocky Mountains plants as C. clavatum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 7 (7 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Phyllary margins ciliate with tiny spreading to recurved spines
var. rivulare
1. Phyllary margins unappendaged or dilated, scarious, and ± lacerate-toothed
→ 2
2. Phyllaries narrowly oblong or linear, often ± subequal, all or most without scarious-dilated margins
var. remotifolium
2. Phyllaries oblong to obovate, often strongly graduated, most or all with dilated, scarious, erose to lacerate-dentate margins
var. odontolepis
1. Plants compact, rounded, moundlike; heads usually not much elevated above leaves
var. compactum
1. Plants usually erect; principal heads usually conspicuously pedunculate
→ 2
2. Corollas white to light purple or rose
var. californicum
2. Corollas deep purple to bright pink or red
→ 3
3. Plants densely white-tomentose; phyllaries persistently white-tomentose (except spines); outer phyllaries usually very long, spreading to reflexed
var. candidissimum
3. Plants variably tomentose, sometimes ± glabrate; phyllaries ± arachnoid to floccose-tomentose, sometimes green and glabrate; outer phyllaries short to long, ascending to spreading or reflexed
→ 4
4. Involucres usually about as long as wide or wider than long; phyllaries densely and persistently arachnoid with fine trichomes connecting tips of adjacent phyllaries
→ 5
4. Involucres usually longer than wide; phyllaries tomentose or glabrate, sparingly or not arachnoid with fine trichomes connecting tips of adjacent phyllaries
→ 6
5. Phyllary apices ± imbricate, the proximal usually shorter than medial and distal, lanceolate to linear-acicular, 0.5–15 mm; co- rollas bright purple
var. occidentale
5. Phyllary apices subequal, all long- acicular, 1.5–3 cm; corollas light to deep reddish purple
var. coulteri
6. Corollas 20–24 mm, deep reddish purple; s Santa Lucia Mountains of San Luis Obispo County, California
var. lucianum
6. Corollas 23–35 mm, bright pink to red; widespread
var. venustum
Source FNA vol. 19, p. 129. FNA vol. 19, p. 137.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. remotifolium var. odontolepis, C. remotifolium var. remotifolium, C. remotifolium var. rivulare
C. occidentale var. californicum, C. occidentale var. candidissimum, C. occidentale var. compactum, C. occidentale var. coulteri, C. occidentale var. lucianum, C. occidentale var. occidentale, C. occidentale var. venustum
Synonyms Carduus remotifolius Carduus occidentalis
Name authority (Hooker) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 655. (1838) (Nuttall) Jepson: Fl. W. Calif., 509. (1901)
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