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fountain thistle

Stems

1–several, erect, loosely arachnoid-tomentose, glandular-pilose;

branches several to many, spreading.

Leaves

blades oblanceolate to oblong or elliptic, 20–70 × 5–16 cm, strongly undulate, shallowly to deeply pinnatifid, lobes coarsely toothed or with triangular secondary lobes, main spines 3–15 mm, abaxial faces more densely tomentose, adaxial densely glandular with short, multicellular trichomes and ± tomentose with fine, arachnoid, non-septate trichomes;

basal often present at flowering, petiolate;

principal cauline well distributed, becoming sessile, gradually reduced distally, bases auriculate-decurrent with broad, spiny-margined wings 3 cm or less;

distal cauline leaves progressively reduced, ± bractlike.

Peduncles

0–7 cm.

Involucres

(green to purple), hemispheric or campanulate, 1.5–3 × 2–5 cm, glabrous or puberulent.

Corollas

white to pinkish or lavender, 14.5–22 mm, tubes 5–8 mm, throats 4.5–9 mm, lobes 3–8 mm;

style tips 2.5–4.5 mm.

Phyllaries

in 6–10 series, imbricate, ovate to lanceolate, abaxial faces without glutinous ridge;

outer and mid bases appressed, short, bodies entire to erose or ciliolate (rarely outer spine-margined), apices spreading to recurved or reflexed, long, dilated, adaxially puberulent, spines 1–6 mm;

apices of inner erect or reflexed, flattened or tipped with short spines.

Heads

few–many, ± nodding, in paniculiform or corymbiform arrays, bracts leafy or much reduced.

Cypselae

brownish, 3.4–5 mm, apical collars tan;

pappi 12–15 mm.

Monocarpic

perennials, 50–220 cm; woody tap-rooted caudices with numerous coarse, fibrous lateral roots, sometimes forming new rosettes from root sprouts.

Cirsium fontinale

Distribution
from USDA
c Calif
[WildflowerSearch map]
[BONAP county map]
Discussion

Varieties 3 (3 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cypselae 4.1–5 mm; phyllary spines 1–2 mm
var. fontinale
1. Cypselae 3.4–4.1 mm; phyllary spines 2–6 mm
→ 2
2. Longer spines of cauline leaves 10–18 mm; most phyllaries without marginal spines
var. campylon
2. Longer spines of cauline leaves 4–7 mm; many outer phyllaries with marginal spines
var. obispoense
Source FNA vol. 19, p. 161.
Parent taxa Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. fontinale var. campylon, C. fontinale var. fontinale, C. fontinale var. obispoense
Synonyms Cnicus fontinalis
Name authority (Greene) Jepson: Fl. W. Calif., 505. (1901)
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