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elk thistle, Evert's thistle, foliose thistle, leafy or foliose or elk thistle, leafy thistle

Habit Biennials or monocarpic perennials, 25–70+ cm; taprooted.
Stems

usually 1, erect, stout, ± fleshy, simple, very leafy, densely villous or tomentose with septate trichomes.

Leaves

blades linear-oblong to oblanceolate (elliptic), 5–20(–25) × 1–4(–7) cm, subentire to dentate or pinnatifid, lobes lance-oblong to triangular, spinulose to spiny-dentate or shallowly lobed, main spines slender, 2–5(–10) mm, abaxial faces often thinly gray- or white-tomentose with felted arachnoid trichomes, ± villous along major veins with septate trichomes, adaxial green, glabrous to thinly arachnoid, often ± villous with septate trichomes;

basal usually present at flowering, spiny winged-petiolate or sessile;

principal cauline well distributed, proximally winged-petiolate, distally sessile, not or only slightly reduced;

distal often narrower than proximal.

Peduncles

0–1 cm.

Involucres

broadly ovoid, 2–2.5 × 1.5–2 cm, green, glabrous to densely villous with septate trichomes on margins.

Corollas

white to pale pink, 21–25 mm, tubes 12–14 mm, throats (very slender, scarcely larger than tubes) 6–7 mm, lobes 3–4 mm;

style tips 2.5–3 mm, short exserted.

Phyllaries

in 4–6 series, imbricate, lanceolate or ovate (outer) to linear-lanceolate (inner), bases appressed, margins of outer entire, abaxial faces without glutinous ridge, apices appressed to ascending, spines straight, slender, 2–3 mm;

apices of inner erect, straight.

Heads

few–many, erect, sessile or subsessile, crowded in dense, woolly, leafy-bracted, subcapitate arrays, closely subtended and overtopped by crowded leafy bracts.

Cypselae

light brown, 4–5.5 mm, apical collars yellow, narrow;

pappi 23–29 mm, exceeding corollas.

2n

= 34.

Cirsium foliosum

Phenology Flowering summer (Jul–Aug).
Habitat Moist soil, grasslands, meadows, edges and openings in boreal forest, subalpine forests and alpine slopes
Elevation 150–2600 m (500–8500 ft)
Distribution
from FNA
WY; AB; BC; NT; YT
[WildflowerSearch map]
[BONAP county map]
Discussion

Cirsium foliosum occurs in the northern Rockies from Wyoming to the Yukon and eastward to the Slave River area in the Northwest Territories and northeastern Alberta. Reports for Alaska are unconfirmed (R. Lipkin, Alaska Natural Heritage Program, pers. comm.). The name Cirsium foliosum has been misapplied to a wide range of plants across the western United States that now are treated as one or another variety of the polymorphic C. scariosum. The only documented occurrences of C. foliosum in the lower 48 states are in the mountains of northern Wyoming. Somewhat similar plants from other mountain areas of the western United States are treated as C. scariosum var. scariosum. During Pleistocene glaciations the ancestors of C. foliosum undoubtedly occupied a more southerly distribution and very likely came into direct contact with ancestral populations of C. scariosum. The observed similarities between C. foliosum and C. scariosum var. scariosum may be a relic of hybridization in that ancient contact zone. On the other hand, the corolla features of C. foliosum suggest that this is a self-pollinating species, perhaps derived from an ancestral population similar to the modern C. scariosum var. scariosum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 19, p. 159.
Parent taxa Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Synonyms Carduus foliosus
Name authority (Hooker) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 654. (1838)
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