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elk thistle, Evert's thistle, foliose thistle, leafy or foliose or elk thistle, leafy thistle

cobwebby thistle, snowy thistle, Venus thistle, western thistle

Habit Biennials or monocarpic perennials, 25–70+ cm; taprooted. Biennials, 5–400 cm; taproots.
Stems

usually 1, erect, stout, ± fleshy, simple, very leafy, densely villous or tomentose with septate trichomes.

usually 1, thinly to densely gray- or white-tomentose, sometimes ± glabrate;

branches few–many, usually from above mid or near base in compact, moundlike dwarf plants, ascending to spreading.

Leaves

blades linear-oblong to oblanceolate (elliptic), 5–20(–25) × 1–4(–7) cm, subentire to dentate or pinnatifid, lobes lance-oblong to triangular, spinulose to spiny-dentate or shallowly lobed, main spines slender, 2–5(–10) mm, abaxial faces often thinly gray- or white-tomentose with felted arachnoid trichomes, ± villous along major veins with septate trichomes, adaxial green, glabrous to thinly arachnoid, often ± villous with septate trichomes;

basal usually present at flowering, spiny winged-petiolate or sessile;

principal cauline well distributed, proximally winged-petiolate, distally sessile, not or only slightly reduced;

distal often narrower than proximal.

blades oblong–elliptic to oblanceolate, 6–40 × 1.5–10+ cm, shallowly to deeply pinnatifid, lobes usually rigidly spreading, undivided or with 1–2 pairs of coarse teeth or lobes, main spines 5–15 mm, both faces gray- to white-tomentose, sometimes ± glabrate or adaxial faces green, thinly arachnoid-tomentose;

basal sometimes present at flowering, petiolate or sessile and bases tapered, spiny-winged;

principal cauline much reduced distally, sessile, bases decurrent or not, as spiny wings;

distal much reduced, linear, ± bractlike.

Peduncles

0–1 cm.

1–30 cm.

Involucres

broadly ovoid, 2–2.5 × 1.5–2 cm, green, glabrous to densely villous with septate trichomes on margins.

ovoid to spheric, 1.5–5 × 1.5–8 cm, arachnoid to ± loosely tomentose, often adjacent phyllaries connected by conspicuous arachnoid trichomes, sometimes glabrous or glabrate.

Corollas

white to pale pink, 21–25 mm, tubes 12–14 mm, throats (very slender, scarcely larger than tubes) 6–7 mm, lobes 3–4 mm;

style tips 2.5–3 mm, short exserted.

white to lavender, pink, rose-purple, or red, 18–40 mm, tubes 8–18 mm, throats 5–7 mm, lobes 5–10 mm;

style tips 4–5 mm.

Phyllaries

in 4–6 series, imbricate, lanceolate or ovate (outer) to linear-lanceolate (inner), bases appressed, margins of outer entire, abaxial faces without glutinous ridge, apices appressed to ascending, spines straight, slender, 2–3 mm;

apices of inner erect, straight.

in 7–10 series, subequal to strongly imbricate, green or stramineous to purple-tinged, linear to narrowly lanceolate, abaxial faces without glutinous ridge;

outer and mid bodies appressed, entire, apices deflexed to spreading or ascending, short-triangular to elongate, linear-acicular, spines spreading to reflexed, 1–10+ mm;

apices of inner erect, often flexuous, flat.

Heads

few–many, erect, sessile or subsessile, crowded in dense, woolly, leafy-bracted, subcapitate arrays, closely subtended and overtopped by crowded leafy bracts.

1–many in loose to tight clusters (barely raised above rosette in dwarf plants).

Cypselae

light brown, 4–5.5 mm, apical collars yellow, narrow;

pappi 23–29 mm, exceeding corollas.

± brown, 5–6 mm, apical collars not differentiated;

pappi 15–30 mm.

2n

= 34.

Cirsium foliosum

Cirsium occidentale

Phenology Flowering summer (Jul–Aug).
Habitat Moist soil, grasslands, meadows, edges and openings in boreal forest, subalpine forests and alpine slopes
Elevation 150–2600 m (500–8500 ft)
Distribution
from FNA
WY; AB; BC; NT; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; NV; OR
[WildflowerSearch map]
[BONAP county map]
Discussion

Cirsium foliosum occurs in the northern Rockies from Wyoming to the Yukon and eastward to the Slave River area in the Northwest Territories and northeastern Alberta. Reports for Alaska are unconfirmed (R. Lipkin, Alaska Natural Heritage Program, pers. comm.). The name Cirsium foliosum has been misapplied to a wide range of plants across the western United States that now are treated as one or another variety of the polymorphic C. scariosum. The only documented occurrences of C. foliosum in the lower 48 states are in the mountains of northern Wyoming. Somewhat similar plants from other mountain areas of the western United States are treated as C. scariosum var. scariosum. During Pleistocene glaciations the ancestors of C. foliosum undoubtedly occupied a more southerly distribution and very likely came into direct contact with ancestral populations of C. scariosum. The observed similarities between C. foliosum and C. scariosum var. scariosum may be a relic of hybridization in that ancient contact zone. On the other hand, the corolla features of C. foliosum suggest that this is a self-pollinating species, perhaps derived from an ancestral population similar to the modern C. scariosum var. scariosum.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 7 (7 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants compact, rounded, moundlike; heads usually not much elevated above leaves
var. compactum
1. Plants usually erect; principal heads usually conspicuously pedunculate
→ 2
2. Corollas white to light purple or rose
var. californicum
2. Corollas deep purple to bright pink or red
→ 3
3. Plants densely white-tomentose; phyllaries persistently white-tomentose (except spines); outer phyllaries usually very long, spreading to reflexed
var. candidissimum
3. Plants variably tomentose, sometimes ± glabrate; phyllaries ± arachnoid to floccose-tomentose, sometimes green and glabrate; outer phyllaries short to long, ascending to spreading or reflexed
→ 4
4. Involucres usually about as long as wide or wider than long; phyllaries densely and persistently arachnoid with fine trichomes connecting tips of adjacent phyllaries
→ 5
4. Involucres usually longer than wide; phyllaries tomentose or glabrate, sparingly or not arachnoid with fine trichomes connecting tips of adjacent phyllaries
→ 6
5. Phyllary apices ± imbricate, the proximal usually shorter than medial and distal, lanceolate to linear-acicular, 0.5–15 mm; co- rollas bright purple
var. occidentale
5. Phyllary apices subequal, all long- acicular, 1.5–3 cm; corollas light to deep reddish purple
var. coulteri
6. Corollas 20–24 mm, deep reddish purple; s Santa Lucia Mountains of San Luis Obispo County, California
var. lucianum
6. Corollas 23–35 mm, bright pink to red; widespread
var. venustum
Source FNA vol. 19, p. 159. FNA vol. 19, p. 137.
Parent taxa Asteraceae > tribe Cardueae > Cirsium Asteraceae > tribe Cardueae > Cirsium
Sibling taxa
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. occidentale, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
C. altissimum, C. andersonii, C. andrewsii, C. arizonicum, C. arvense, C. barnebyi, C. brevifolium, C. brevistylum, C. canescens, C. carolinianum, C. ciliolatum, C. clavatum, C. crassicaule, C. cymosum, C. discolor, C. douglasii, C. drummondii, C. eatonii, C. edule, C. engelmannii, C. flodmanii, C. foliosum, C. fontinale, C. grahamii, C. helenioides, C. hookerianum, C. horridulum, C. hydrophilum, C. inamoenum, C. joannae, C. kamtschaticum, C. lecontei, C. longistylum, C. mohavense, C. muticum, C. neomexicanum, C. nuttallii, C. ochrocentrum, C. ownbeyi, C. palustre, C. parryi, C. perplexans, C. pitcheri, C. praeteriens, C. pulcherrimum, C. pumilum, C. quercetorum, C. remotifolium, C. repandum, C. rhothophilum, C. rydbergii, C. scariosum, C. texanum, C. tracyi, C. turneri, C. undulatum, C. vinaceum, C. virginianum, C. vulgare, C. wheeleri, C. wrightii
Subordinate taxa
C. occidentale var. californicum, C. occidentale var. candidissimum, C. occidentale var. compactum, C. occidentale var. coulteri, C. occidentale var. lucianum, C. occidentale var. occidentale, C. occidentale var. venustum
Synonyms Carduus foliosus Carduus occidentalis
Name authority (Hooker) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 654. (1838) (Nuttall) Jepson: Fl. W. Calif., 509. (1901)
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