Cirsium foliosum |
Cirsium occidentale |
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elk thistle, Evert's thistle, foliose thistle, leafy or foliose or elk thistle, leafy thistle |
cobwebby thistle, snowy thistle, Venus thistle, western thistle |
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Habit | Biennials or monocarpic perennials, 25–70+ cm; taprooted. | Biennials, 5–400 cm; taproots. | ||||||||||||||||||||||||
Stems | usually 1, erect, stout, ± fleshy, simple, very leafy, densely villous or tomentose with septate trichomes. |
usually 1, thinly to densely gray- or white-tomentose, sometimes ± glabrate; branches few–many, usually from above mid or near base in compact, moundlike dwarf plants, ascending to spreading. |
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Leaves | blades linear-oblong to oblanceolate (elliptic), 5–20(–25) × 1–4(–7) cm, subentire to dentate or pinnatifid, lobes lance-oblong to triangular, spinulose to spiny-dentate or shallowly lobed, main spines slender, 2–5(–10) mm, abaxial faces often thinly gray- or white-tomentose with felted arachnoid trichomes, ± villous along major veins with septate trichomes, adaxial green, glabrous to thinly arachnoid, often ± villous with septate trichomes; basal usually present at flowering, spiny winged-petiolate or sessile; principal cauline well distributed, proximally winged-petiolate, distally sessile, not or only slightly reduced; distal often narrower than proximal. |
blades oblong–elliptic to oblanceolate, 6–40 × 1.5–10+ cm, shallowly to deeply pinnatifid, lobes usually rigidly spreading, undivided or with 1–2 pairs of coarse teeth or lobes, main spines 5–15 mm, both faces gray- to white-tomentose, sometimes ± glabrate or adaxial faces green, thinly arachnoid-tomentose; basal sometimes present at flowering, petiolate or sessile and bases tapered, spiny-winged; principal cauline much reduced distally, sessile, bases decurrent or not, as spiny wings; distal much reduced, linear, ± bractlike. |
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Peduncles | 0–1 cm. |
1–30 cm. |
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Involucres | broadly ovoid, 2–2.5 × 1.5–2 cm, green, glabrous to densely villous with septate trichomes on margins. |
ovoid to spheric, 1.5–5 × 1.5–8 cm, arachnoid to ± loosely tomentose, often adjacent phyllaries connected by conspicuous arachnoid trichomes, sometimes glabrous or glabrate. |
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Corollas | white to pale pink, 21–25 mm, tubes 12–14 mm, throats (very slender, scarcely larger than tubes) 6–7 mm, lobes 3–4 mm; style tips 2.5–3 mm, short exserted. |
white to lavender, pink, rose-purple, or red, 18–40 mm, tubes 8–18 mm, throats 5–7 mm, lobes 5–10 mm; style tips 4–5 mm. |
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Phyllaries | in 4–6 series, imbricate, lanceolate or ovate (outer) to linear-lanceolate (inner), bases appressed, margins of outer entire, abaxial faces without glutinous ridge, apices appressed to ascending, spines straight, slender, 2–3 mm; apices of inner erect, straight. |
in 7–10 series, subequal to strongly imbricate, green or stramineous to purple-tinged, linear to narrowly lanceolate, abaxial faces without glutinous ridge; outer and mid bodies appressed, entire, apices deflexed to spreading or ascending, short-triangular to elongate, linear-acicular, spines spreading to reflexed, 1–10+ mm; apices of inner erect, often flexuous, flat. |
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Heads | few–many, erect, sessile or subsessile, crowded in dense, woolly, leafy-bracted, subcapitate arrays, closely subtended and overtopped by crowded leafy bracts. |
1–many in loose to tight clusters (barely raised above rosette in dwarf plants). |
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Cypselae | light brown, 4–5.5 mm, apical collars yellow, narrow; pappi 23–29 mm, exceeding corollas. |
± brown, 5–6 mm, apical collars not differentiated; pappi 15–30 mm. |
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2n | = 34. |
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Cirsium foliosum |
Cirsium occidentale |
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Phenology | Flowering summer (Jul–Aug). | |||||||||||||||||||||||||
Habitat | Moist soil, grasslands, meadows, edges and openings in boreal forest, subalpine forests and alpine slopes | |||||||||||||||||||||||||
Elevation | 150–2600 m (500–8500 ft) | |||||||||||||||||||||||||
Distribution |
WY; AB; BC; NT; YT
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CA; NV; OR
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Discussion | Cirsium foliosum occurs in the northern Rockies from Wyoming to the Yukon and eastward to the Slave River area in the Northwest Territories and northeastern Alberta. Reports for Alaska are unconfirmed (R. Lipkin, Alaska Natural Heritage Program, pers. comm.). The name Cirsium foliosum has been misapplied to a wide range of plants across the western United States that now are treated as one or another variety of the polymorphic C. scariosum. The only documented occurrences of C. foliosum in the lower 48 states are in the mountains of northern Wyoming. Somewhat similar plants from other mountain areas of the western United States are treated as C. scariosum var. scariosum. During Pleistocene glaciations the ancestors of C. foliosum undoubtedly occupied a more southerly distribution and very likely came into direct contact with ancestral populations of C. scariosum. The observed similarities between C. foliosum and C. scariosum var. scariosum may be a relic of hybridization in that ancient contact zone. On the other hand, the corolla features of C. foliosum suggest that this is a self-pollinating species, perhaps derived from an ancestral population similar to the modern C. scariosum var. scariosum. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 7 (7 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 19, p. 159. | FNA vol. 19, p. 137. | ||||||||||||||||||||||||
Parent taxa | Asteraceae > tribe Cardueae > Cirsium | Asteraceae > tribe Cardueae > Cirsium | ||||||||||||||||||||||||
Sibling taxa | ||||||||||||||||||||||||||
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Synonyms | Carduus foliosus | Carduus occidentalis | ||||||||||||||||||||||||
Name authority | (Hooker) de Candolle: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 6: 654. (1838) | (Nuttall) Jepson: Fl. W. Calif., 509. (1901) | ||||||||||||||||||||||||
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