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goldenaster

highlands goldenaster

Habit Biennials, perennials, or subshrubs, 20–110(–150) cm, hairs usually soft, flexible, flagelliform (1–5 large, short basal cells, abruptly changing to thinner, elongate distal cells); taprooted or fibrous-rooted. Perennials (weak or monocarpic), (30–)60–110 cm; taprooted (new rosettes from bases of previous season’s stems or as side branches of primary unbolted rosettes).
Stems

erect, ascending, procumbent or decumbent, simple or branched, glabrous or woolly, arachnoid (in C. mariana), often densely stipitate-glandular, especially distally.

1–8(–20), usually ascending to erect, moderately spreading near base, rarely decumbent, frequently branched (densely leafy), sparsely lanate, stipitate-glandular distally.

Leaves

basal (in rosettes) and cauline; alternate;

sessile;

basal blades 1-nerved (net-veined), spatulate to oblanceolate (bases petiole-like), margins entire or apically dentate-serrate, faces usually sparsely to densely woolly (at least in young rosettes) or piloso-sericeous, rarely arachnoid;

cauline blades linear, lanceolate, elliptic, or ovate (bases sometimes ± clasping), margins entire or dentate, sometimes coarsely ciliate, faces glabrous or densely woolly, pilose, or arachnoid, or stipitate-glandular.

basal blades usually spatulate to oblanceolate, rarely obovate, (35–)55–120(–150) × 15–20(–30) mm, bases narrowly attenuate-cuneate, margins entire or apically serrate-denticulate, faces loosely long-lanate, obscurely stipitate-glandular, viscid;

cauline sessile, blades usually oblong to oblong-elliptic, sometimes ovate to lanceolate, bases usually slightly auriculate-clasping, truncate to rounded, margins undulate, some cilia 2–3 mm, faces moderately woolly-lanulate, stipitate-glandular;

mid to distal leaves ascending, often appressed, apices usually obtuse, sometimes subacute, mucronate or mucronulate, faces stipitate-glandular, distalmost glabrate to sparsely woolly-pilose, densely stipitae-glandular.

Involucres

campanulate, (5–12 ×) 5.5–14 mm.

(yellow-green in bud) cylindro-campanulate, 6–8.5 mm.

Receptacles

slightly convex, shallowly pitted, epaleate.

Ray florets

9–36, pistillate, fertile;

corollas yellow.

16–22;

laminae 8–9.5 × 1–2 mm.

Disc florets

25–90, bisexual, fertile;

corollas yellow, ampliate, tubes ± equaling distally dilated throats, lobes 5, erect, triangular;

style-branch appendages triangular.

20–40;

corollas 4.5–5.5 mm, lobes 0.6–1 mm.

Phyllaries

30–55 in 3–5 series, 1-nerved, (weakly to strongly keeled proximally, flat to slightly convex distally), linear-lanceolate to oblanceolate, unequal, chartaceous to membranous proximally, membranous or foliaceous distally, margins scarious proximally, faces glabrous or hairy, sometimes stipitate-glandular or gland-dotted.

in 5–6 series, erect, linear-lanceolate, unequal, 0.7–1.1 mm wide, apices usually acute-acuminate, sometimes acute-aristate, sometimes inner obtuse to rounded, densely stipitate-glandular, viscid.

Heads

in corymbiform, subumbelliform, or paniculiform arrays (peduncles nearly naked to leafy-bracteate).

10–50, in compact corymbiform to paniculiform arrays (height usually less than 1/5 plants, branches slender, ascending, stipitate-glandular, sweet camphor smelling).

Cypselae

(stramineous) obconic, compressed, usually smooth or shallowly 1–10-ribbed, sometimes 2–10-ridged (ridges yellow to red-brown, clavate, translucent), faces sparsely to moderately strigose;

pappi persistent, in (2–)3(–4) series, outer of linear to narrowly triangular scales 0.4–1.4 mm, inner 1–2(–3) series of 20–40 whitish to stramineous, thin, barbellate, apically attenuate or clavate bristles.

2–2.7 mm, without ridges, shallowly ribbed or smooth, moderately long-strigose;

pappi in 3–4 series, outer of linear scales 0.5–1.5 mm, inner of 30–35 bristles 5–6 mm, inner moderately clavate.

x

= 5, 4, 9.

2n

= 10.

Chrysopsis

Chrysopsis highlandsensis

Phenology Flowering late Oct–mid Dec.
Habitat Sand pine scrub, scrubby flatwoods
Elevation 20–50 m (100–200 ft)
Distribution
from USDA
se United States
[BONAP county map]
from FNA
FL
[BONAP county map]
Discussion

Species 11 (11 in the flora).

All species of Chrysopsis are native to Florida; two species occur also north and west of the state. As circumscribed here, the genus includes the North American goldenasters that bear flagelliform hairs with bases of large cells. Historically, Chrysopsis included most species treated here in Heterotheca and all species treated here in Pityopsis (J. C. Semple 1981, 1996, literature cited therein). The two annual species of Bradburia are excluded, although G. L. Nesom (1991) included them in Chrysopsis. The cytotaxonomy of Chrysopsis has been studied in detail with the karyotypes of nearly all species illustrated, the allopolypoid origins of the C. gossypina complex documented, and the cytogeography of the four ploidy levels in C. mariana described (Semple 1977; Semple et al. 1980; Semple and C. C. Chinnappa 1980, 1980b, 1984, 1986). Semple and J. L. A. Hood (2005) described the pappi of the genera of subtribe Chrysopsidinae. Pappus traits of species within Chrysopsis are similar but differ from those of Bradburia, Heterotheca, and Pityopsis. Most species of Chrysopsis are restricted to limited habitats within Florida; some are rare or threatened. Different morphotypes of C. gossypina, a variable allopolyploid, may be superficially similar to one or another of the distinct diploid species (Semple 1981).

Diplogon Rafinesque is a rejected name.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Of conservation concern.

Chrysopsis highlandsensis is known from Highlands, southern Polk, and northern Glades counties.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Cypselae 2–10-ridged (ridges yellow to red-brown, translucent); phyllaries glabrous or densely pilose, sometimes stipitate-glandular
→ 2
1. Cypselae without yellow to red-brown, translucent ridges; phyllaries moderately to densely glandular
→ 6
2. Phyllary apices spreading to reflexed, twisted, usually long-attenuate (some n Florida collections acute, but leaves typical), eglandular; cypselae 6–10-ridged
C. subulata
2. Phyllary apices usually appressed, acute to acuminate, sometimes stipitate-glandular; cypselae 2–6-ridged
→ 3
3. Phyllaries 1.5–2.5 mm wide
C. latisquamea
3. Phyllaries 0.5–1(–1.3) mm wide
→ 4
4. Margins of distal leaves piloso-ciliate
C. gossypina
4. Margins of distal leaves not piloso-ciliate
→ 5
5. Stems erect; leaf blades mostly linear (sometimes twisted), proximal to mid cauline faces glabrous; panhandle and peninsular Florida
C. linearifolia
5. Stems procumbent; leaf blades linear-elliptic to oblong; proximal to mid cauline faces sometimes sparsely pilose
C. gossypina
6. Distal leaves (and stems) silky-sericeous
C. mariana
6. Distal leaves stipitate-glandular, hirsute, woolly, lanulate, or sericeous-tomentose, not silky-sericeous.
→ 7
7. Phyllary apices spreading to reflexed, attenuate to long-attenuate; c, w Florida panhandle
→ 8
7. Phyllary apices erect, acute, acuminate, or obtuse; peninsular and e panhandle Florida
→ 9
8. Stems decumbent to ascending or erect; heads in corymbiform to paniculiform arrays; coastal dunes (see also forms of C. gossypina subsp. gossypina)
C. godfreyi
8. Stems erect; heads in paniculiform arrays; inland in disturbed habitats
C. lanuginosa
9. Mid cauline leaf faces woolly or tomentose; involucres yellow-green in bud
→ 10
9. Mid cauline leaf faces usually sparsely hirsute, rarely woolly, stipitate-glandular; involucres green in bud
→ 11
10. Mid cauline leaf blades obovate or oblanceolate, bases cuneate to slightly auriculate-clasping, marginal cilia rarely more than 1 mm, faces appressed-tomentose; distal blade faces appressed-tomentose, sparsely stipitate-glandular (glands hidden by hair)
C. floridana
10. Mid cauline leaf blades oblong, oblong-elliptic, sometimes ovate to lanceolate, bases truncate to rounded, some marginal cilia 2–3 mm, faces woolly-lanulate; distal blade faces glabrate to sparsely woolly-pilose, densely stipitate-glandular
C. highlandsensis
11. Biennials; stems erect, simple, tips erect to ascending before heads form; mid leaf faces glabrate to sparsely strigillose, densely short-stipitate-glandular (larger glandular hairs 0.01–0.3 mm); heads in compact to lax, corymbiform arrays; peninsu-lar and e panhandle Florida
C. scabrella
11. Perennials (sometimes monocarpic); stems ascending to erect, tips usually nodding before heads form; mid leaf faces moderately hirsute, densely stipitate-glandular, viscid (larger glandular hairs 0.3–0.9 mm); heads in compact to moderately open, corymbiform to paniculiform arrays; mostly Lake Wales Ridge and Atlantic Ridge, c and se Florida
C. delaneyi
Source FNA vol. 20, p. 213. Author: John C. Semple. FNA vol. 20, p. 216.
Parent taxa Asteraceae > tribe Astereae Asteraceae > tribe Astereae > Chrysopsis
Sibling taxa
C. delaneyi, C. floridana, C. godfreyi, C. gossypina, C. lanuginosa, C. latisquamea, C. linearifolia, C. mariana, C. scabrella, C. subulata
Subordinate taxa
C. delaneyi, C. floridana, C. godfreyi, C. gossypina, C. highlandsensis, C. lanuginosa, C. latisquamea, C. linearifolia, C. mariana, C. scabrella, C. subulata
Synonyms Inula, Diplopappus, Heterotheca section C.
Name authority (Nuttall) Elliott: Sketch Bot. S. Carolina 2: 333. (1823) Delaney & Wunderlin: Bot. Explor. 2: 2, figs. 1–3, 5, 7, 9, 11. (2002)
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