Flora of North America species comparison

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Cheilanthes covillei

Coville's lip fern
Stems

short-creeping, usually 2–4 mm diam.;

scales usually uniformly dark brown to black or rarely with narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent.
Leaves

clustered, 5–30 cm;

vernation noncircinate.
Petiole

dark brown, rounded adaxially.
Blade

lanceolate to ovate-deltate, 3–4-pinnate at base, 1.5–5 cm wide;

rachis rounded adaxially, somewhat scaly, not pubescent.
Ultimate segments

round to oblong, beadlike, the largest 1–3 mm, abaxially glabrous or with a few small scales near base, adaxially glabrous.
Pinnae

not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous (or somewhat scaly) adaxially.
False indusia

marginal, weakly differentiated, 0.05–0.25 mm wide.
Sori

± continuous around segment margins.
Sporangia

containing 64 spores.
Costae

green adaxially for most of length;

abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the largest 0.4–1.5 mm wide, strongly imbricate, usually concealing ultimate segments, ciliate only on basal lobes.
2n

= 60.

Cheilanthes covillei
Phenology Sporulating late spring–fall.
Habitat Rocky slopes, cliffs, and ledges, usually on igneous substrates
Elevation 100–2500 m (300–8200 ft)
Distribution
from FNA
AZ; CA; NV; UT; Mexico in Baja California
[WildflowerSearch map]
Discussion

Cheilanthes covillei can be difficult to distinguish from the closely related C. intertexta and C. clevelandii; it differs from these two species in having glabrous blades and costal scales ciliate only on the basal lobes. Cheilanthes covillei is occasionally misidentified as C. fendleri because the cilia of the scales are often obscure; it is distinguished from the latter species by having rigid, dark brown stem scales that are strongly appressed. Cheilanthes covillei hybridizes with C. parryi and C. newberryi to form rare, sterile diploids known as C. × parishii Davenport and C. × fibrillosa (Davenport) Davenport ex Underwood, respectively. A third sterile diploid hybrid with C. fendleri has recently been discovered in central Arizona (M. D. Windham, unpublished).

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Source FNA vol. 2.
Parent taxa Pteridaceae > Cheilanthes
Sibling taxa
C. aemula, C. alabamensis, C. arizonica, C. bonariensis, C. clevelandii, C. cooperae, C. eatonii, C. feei, C. fendleri, C. gracillima, C. horridula, C. intertexta, C. kaulfussii, C. lanosa, C. lendigera, C. leucopoda, C. lindheimeri, C. microphylla, C. newberryi, C. parryi, C. pringlei, C. tomentosa, C. villosa, C. viscida, C. wootonii, C. wrightii, C. yavapensis
Synonyms Myriopteris covillei
Name authority Maxon: Proc. Biol. Soc. Wash. 31: 147. (1918)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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