Cheilanthes covillei |
Cheilanthes fendleri |
|
---|---|---|
Coville's lip fern |
Fendler's lip fern |
|
Stems | short-creeping, usually 2–4 mm diam.; scales usually uniformly dark brown to black or rarely with narrow, light brown margins, linear-lanceolate, straight to slightly contorted, strongly appressed, persistent. |
long-creeping, 1–3 mm diam.; scales mostly uniformly brown, ovate-lanceolate, straight to slightly contorted, loosely appressed, often deciduous on older portions of stem. |
Leaves | clustered, 5–30 cm; vernation noncircinate. |
scattered, 7–30 cm; vernation noncircinate. |
Petiole | dark brown, rounded adaxially. |
usually dark brown, rounded adaxially. |
Blade | lanceolate to ovate-deltate, 3–4-pinnate at base, 1.5–5 cm wide; rachis rounded adaxially, somewhat scaly, not pubescent. |
lanceolate to ovate-deltate, 3–4-pinnate at base, 1.5–5 cm wide; rachis rounded adaxially, somewhat scaly, not pubescent. |
Ultimate segments | round to oblong, beadlike, the largest 1–3 mm, abaxially glabrous or with a few small scales near base, adaxially glabrous. |
round to oblong, beadlike, the largest 1.5–3 mm, abaxially glabrous or with a few small scales near base, adaxially glabrous. |
Pinnae | not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous (or somewhat scaly) adaxially. |
not articulate, dark color of stalk continuing into pinna base, basal pair not conspicuously larger than adjacent pair, usually equilateral, appearing glabrous adaxially. |
False indusia | marginal, weakly differentiated, 0.05–0.25 mm wide. |
marginal, weakly differentiated, 0.05–0.25 mm wide. |
Sori | ± continuous around segment margins. |
± continuous around segment margins. |
Sporangia | containing 64 spores. |
containing 64 spores. |
Costae | green adaxially for most of length; abaxial scales multiseriate, ovate-lanceolate, deeply cordate at base, with overlapping basal lobes, conspicuous, the largest 0.4–1.5 mm wide, strongly imbricate, usually concealing ultimate segments, ciliate only on basal lobes. |
green adaxially for most of length; abaxial scales multiseriate, lanceolate-ovate, truncate or subcordate at base, without overlapping basal lobes, conspicuous, the largest 0.4–1.2 mm wide, strongly imbricate, often concealing ultimate segments, entire to denticulate, not ciliate. |
2n | = 60. |
= 60. |
Cheilanthes covillei |
Cheilanthes fendleri |
|
Phenology | Sporulating late spring–fall. | Sporulating summer–fall. |
Habitat | Rocky slopes, cliffs, and ledges, usually on igneous substrates | Rocky slopes and ledges, found on a variety of acidic and mildly basic substrates |
Elevation | 100–2500 m (300–8200 ft) | 1200–3100 m (3900–10200 ft) |
Distribution |
AZ; CA; NV; UT; Mexico in Baja California
|
AZ; CO; NM; TX; n Mexico |
Discussion | Cheilanthes covillei can be difficult to distinguish from the closely related C. intertexta and C. clevelandii; it differs from these two species in having glabrous blades and costal scales ciliate only on the basal lobes. Cheilanthes covillei is occasionally misidentified as C. fendleri because the cilia of the scales are often obscure; it is distinguished from the latter species by having rigid, dark brown stem scales that are strongly appressed. Cheilanthes covillei hybridizes with C. parryi and C. newberryi to form rare, sterile diploids known as C. × parishii Davenport and C. × fibrillosa (Davenport) Davenport ex Underwood, respectively. A third sterile diploid hybrid with C. fendleri has recently been discovered in central Arizona (M. D. Windham, unpublished). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Young, sterile plants of Cheilanthes fendleri are occasionally misidentified as C. pringlei; they are distinguished from that species by having rachises that are rounded (not grooved) adaxially. Cheilanthes fendleri apparently hybridizes with both C. wootonii (T. Reeves 1979) and C. covillei (M. D. Windham, unpublished). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | Pteridaceae > Cheilanthes | Pteridaceae > Cheilanthes |
Sibling taxa | ||
Synonyms | Myriopteris covillei | Myriopteris fendleri |
Name authority | Maxon: Proc. Biol. Soc. Wash. 31: 147. (1918) | Hooker: Sp. Fil. 2: 103, plate 107b. (1852) |
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