Centaurea melitensis |
Centaurea nigra |
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croix de malte, Maltese knapweed, Maltese star thistle or centaury, Maltese star-thistle, Napa thistle, tocalote |
black knapweed, centaurée noire, common knapweed, hardheads, lesser knapweed |
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Habit | Annuals, 10–100 cm, herbage loosely gray-tomentose and villous with jointed multicellular hairs, sometimes minutely scabrous, minutely resin-gland-dotted. | Perennials, 30–150 cm. |
Stems | 1–few, few–many branched distally. |
1–few, erect or ascending, openly branched distally, villous to scabrous with septate hairs and loosely tomentose, ± glabrate. |
Leaves | basal and proximal cauline petiolate or tapering to base, usually absent at anthesis, blades oblong to oblanceolate, 2–15 cm, margins entire to dentate or pinnately lobed; cauline long-decurrent, blades linear to oblong or oblanceolate, 1–5 cm, entire or dentate. |
basal and proximal cauline petiolate, blades oblanceolate or elliptic, 5–25 cm, margins entire or shallowly dentate to irregularly pinnately lobed; distal cauline sessile, not decurrent, gradually smaller, blades linear to lanceolate, entire or dentate. |
Involucres | ovoid, 10–15 mm, loosely cobwebby-tomentose or becoming glabrous. |
ovoid to campanulate or hemispheric, 15–18 mm, usually ± as wide as high. |
Florets | many; corollas yellow, those of sterile florets 10–12 mm, slender, inconspicuous, those of fertile florets 10–12 mm. |
40–100+, all fertile; corollas purple (rarely white), 15–18 mm. |
Inner phyllaries | appendages entire, acute or spine-tipped. |
tips truncate, irregularly dentate or lobed. |
Heads | disciform, 1–few at branch tips, borne singly or in open leafy corymbiform arrays, sometimes clustered in distal axils, sessile or pedunculate. |
discoid, in few-headed corymbiform arrays, borne on leafy-bracted peduncles. |
Cypselae | dull white or light brown, ca. 2.5 mm, finely hairy; pappi of many white, unequal, stiff bristles 2.5–3 mm. |
tan, 2.5–3 mm, finely hairy; pappi of many blackish, unequal, sometimes deciduous bristles 0.5–1 mm. |
Principal | phyllaries: bodies ± stramineous, ovate, appendages purplish, spiny-fringed at base, each tipped by slender spine 5–10 mm. |
phyllaries: bodies lanceolate to ovate, loosely tomentose or glabrous, bases usually ± concealed by expanded appendages, appendages erect, overlapping, dark brown to black, flat, margins pectinately dissected into numerous wiry lobes. |
2n | = 24. |
= 22, 44. |
Centaurea melitensis |
Centaurea nigra |
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Phenology | Flowering mostly spring–summer (Apr–Jul). | Flowering summer–fall (Jun–Oct). |
Habitat | Roadsides, fields, pine-oak woodlands, chaparral, agricultural areas | Roadsides, fields, clearings, waste areas |
Elevation | 0–1500 m (0–4900 ft) | 0–300 m (0–1000 ft) |
Distribution |
AL; AZ; CA; GA; ID; IL; MA; MO; MS; NJ; NM; NV; OR; PA; TX; UT; WA; WI; BC; Mexico (Baja California); Europe; Asia; Africa [Widely introduced]
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CA; CT; DE; IA; ID; IL; KY; MA; MD; ME; MI; MO; MT; NH; NJ; NY; OH; OR; PA; RI; VA; VT; WA; WI; WV; BC; NB; NF; NS; ON; PE; QC; SPM; Europe [Introduced in North America]
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Discussion | Centaurea melitensis is native to the Mediterranean region. It is listed as a noxious weed in New Mexico. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Black knapweed is listed as a noxious weed in Colorado and Washington. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 19, p. 193. | FNA vol. 19, p. 187. |
Parent taxa | Asteraceae > tribe Cardueae > Centaurea | Asteraceae > tribe Cardueae > Centaurea |
Sibling taxa | ||
Synonyms | C. jacea subsp. nigra, C. nemoralis | |
Name authority | Linnaeus: Sp. Pl. 2: 917. (1753) | Linnaeus: Sp. Pl. 2: 911. (1753) |
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