Cardamine nuttallii |
Cardamine pattersonii |
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beautiful bitter-cress, beautiful bittercres, Nuttall's bittercress, Nuttall's toothwort, palmate toothwort, slender toothwort |
Saddle Mountain bitter-cress |
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Habit | Perennials; glabrous or sparsely pubescent. | Annuals or perennials; (short-lived); glabrous throughout. |
Rhizomes | (tuberiform, fragile), with ovoid to oblong or cylindrical nodal swellings, slender, 2–5 mm diam., (fleshy). |
cylindrical, slender, 0.5–1.5 mm diam. |
Stems | erect, unbranched, 0.5–2(–3) dm, glabrous or sparsely pubescent distally. |
(simple from base), erect, (not flexuous), unbranched or branched basally, 0.6–3 dm. |
Basal leaves | rosulate or not, 3 or 5-foliolate, 1–6 cm, leaflets petiolulate; petiole 0.4–2.5 cm; lateral leaflets similar to terminal, considerably smaller; terminal leaflet (petiolule 0.1–0.4 cm), blade obovate to orbicular or subcordate, 0.3–1.5(–2.0) cm × 2.5–16(–18) mm, base obtuse to rounded or cordate, margins entire or dentate to slightly sinuately lobed. |
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Rhizomal leaves | simple or 3 (or 5)-foliolate, (3–)4–20(–25) cm, leaflets petiolulate or subsessile; petiole (2–)3–18(–21) cm; lateral leaflets (when present) petiolulate to subsessile, blade similar to terminal, sometimes smaller; terminal leaflet (subsessile or petiolule 0.2–3 cm), blade (simple leaf or terminal leaflet) reniform to suborbicular or ovate to oblong, (0.9–)1.3–4(–5.2) cm × (8–)12–50(–70) mm, base cordate to obtuse, margins crenate, dentate, or 5–7-lobed, (apiculae terminating teeth or lobes, surfaces glabrous). |
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Cauline leaves | 1–3, 3 (or 5)-foliolate, (appearing palmate), petiolate, leaflets petiolulate or sessile; petiole (0.2–)0.5–2(–3) cm, base not auriculate; lateral leaflets sessile, blade similar to terminal, smaller; terminal leaflet petiolulate or sessile, blade broadly ovate to oblong or linear, (0.5–)1–3.5(–6) cm, margins usually entire or dentate, rarely lobed. |
2–4, 3 or 5-foliolate, blade similar to basal, gradually reduced in size as bract, distalmost ones subtending pedicels of flowers (usually simple); leaflet blade or bract linear to linear-oblanceolate, 0.2–0.8 cm. |
Racemes | ebracteate. |
bracteate throughout. |
Flowers | sepals oblong, 3.5–5 × 1.5–2 mm, lateral pair saccate basally; petals usually purple to pale pink, rarely white, obovate, 10–15 × 4–7.5 mm, (not clawed, apex rounded); filaments: median pairs 5–8 mm, lateral pair 3.5–5 mm; anthers oblong, 1.5–2 mm. |
sepals oblong, (2–)3 × 0.7(–1) mm, lateral pair not saccate basally; petals purple or pink, obovate, 6–9 × 3–4 mm, (not clawed); filaments: median pairs 3–3.5 mm, lateral pair 2–2.5 mm; anthers oblong, 0.7–1 mm. |
Fruiting pedicels | ascending to divaricate, 10–35 mm. |
divaricate-ascending, (10–)15–30(–45) mm. |
Fruits | linear, 2.5–5.6 cm × 2–2.3 mm; ovules 8–16 per ovary; style 4–8 mm. |
linear, (torulose), 2–3 cm × 1–1.5 mm; ovules 14–20 per ovary; style 2–4 mm. |
Seeds | dark brown, oblong, 2–2.5 × 1.4–1.6 mm. |
brown, oblong to ovoid, 1.7–2.2 × 1–1.5 mm, (winged distally). |
Cardamine nuttallii |
Cardamine pattersonii |
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Phenology | Flowering Mar–May. | Flowering May–Jun. |
Habitat | Open pine forests, damp woods, shaded bottomlands, mossy slopes, streamsides, shaded and moist hillsides | Moist mossy cliffs, rocky slopes, mossy banks |
Elevation | 150-1000 m (500-3300 ft) | 800-900 m (2600-3000 ft) |
Distribution |
CA; OR; WA; BC
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OR
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Discussion | The infraspecific taxonomy of Cardamine nuttallii has been based almost entirely on the division and margin of rhizomal leaves. The treatments by O. E. Schulz (1903), L. E. Detling (1937), and R. C. Rollins (1993), though utilizing the same characters, varied considerably, especially in the application of names to varieties. The absence of rhizomal leaves on most specimens makes varietal determination an almost impossible task. Furthermore, leaf morphology is so highly variable that it is not useful for formally recognizing some of the other variants in the species. We therefore prefer to not subdivide the species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Cardamine pattersonii is known from Saddle Mountain and Onion Peak in Clastop County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 478. | FNA vol. 7, p. 480. |
Parent taxa | Brassicaceae > tribe Cardamineae > Cardamine | Brassicaceae > tribe Cardamineae > Cardamine |
Sibling taxa | ||
Synonyms | Dentaria tenella, C. californica var. gemmata, C. gemmata, C. nuttallii var. covilleana, C. nuttallii var. dissecta, C. nuttallii var. gemmata, C. nuttallii var. pulcherrima, C. pulcherrima, C. pulcherrima var. tenella, C. quercetorum, C. tenella var. covilleana, C. tenella var. dissecta, C. tenella var. quercetorum, Dentaria gemmata, Dentaria macrocarpa, Dentaria macrocarpa var. pulcherrima, Dentaria quercetorum, Dentaria tenella var. palmata, Dentaria tenella var. pulcherrima, Dentaria tenella var. quercetorum | |
Name authority | Greene: Bull. Calif. Acad. Sci. 2: 389. (1887) | L. F. Henderson: Rhodora 32: 25. (1930) |
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