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dwarf braya, maccall's dwarf braya, mccalla's dwarf braya

northern-rockcress

Habit Perennials; (sometimes pulvinate, caudex simple or many-branched); scapose or not; usually pubescent or pilose, sometimes glabrous, trichomes short-stalked, forked, subdendritic, or submalpighiaceous, mixed with simple ones (rarely exclusively).
Stems

ascending to erect, often unbranched, 0.4–2.3 dm, moderately pubescent.

erect to decumbent or ascending, unbranched or branched.

Leaves

blade margins entire or, sometimes, sinuate-dentate, surfaces glabrescent to moderately pubescent.

basal and, sometimes, cauline;

petiolate or sessile;

basal rosulate, petiolate, blade margins entire, sinuate, dentate, or, rarely, pinnately lobed;

cauline usually absent, rarely few present, (sub)sessile, blade margins usually entire, rarely dentate or pinnately lobed.

Racemes

(corymbose, sometimes bracteate basally or throughout), elongated or not in fruit.

Flowers

petals white, (4.4–)4.9–6.7(–7.2) × (2.1–)2.3–4(–4.2) mm.

sepals [sometimes persistent], oblong [ovate], lateral pair not saccate basally (sometimes slightly so in B. humilis and B. linearis);

petals white, pink, or purple [rarely pale yellow], obovate, oblanceolate, or spatulate, (slightly to much longer than sepals), claw distinct or not, (shorter than sepal, apex obtuse or rounded);

stamens tetradynamous;

filaments dilated or not basally;

anthers ovate or oblong, (apex usually obtuse, sometimes apiculate);

nectar glands (4), lateral, 1 on each side of lateral stamen.

Fruiting pedicels

erect, divaricate, or ascending, slender (much narrower than fruit).

Fruits

often abortive, somewhat torulose, 0.7–1.2 mm wide when fully developed;

septum not fenestrate or split longitudinally.

siliques or silicles, sessile, linear, oblong, cylindrical, oval-elliptic, ovoid, lanceoloid, lanceoloid-subulate, or globose, smooth or torulose, terete or slightly latiseptate;

valves each often with prominent midvein, glabrous or pubescent;

replum rounded;

septum complete, (membranous, translucent);

ovules (5–)14–44 per ovary;

stigma capitate, entire or slightly 2-lobed.

Seeds

plump, not winged, oblong or ovoid;

seed coat (minutely reticulate), not mucilaginous when wetted;

cotyledons incumbent.

x

= 7.

2n

= 28.

Braya humilis subsp. maccallae

Braya

Phenology Flowering May–Jul.
Habitat Sandy gravelly riverbanks and floodplains, sometimes on slopes and glacial moraines
Elevation 1500-3000 m (4900-9800 ft)
Distribution
from FNA
AB; BC
[BONAP county map]
from USDA
North America; n Europe; Asia
[BONAP county map]
Discussion

Subspecies maccallae differs from subspp. elesmerensis and humilis in having leaf margins mostly entire, flowers exceptionally large, and a high percentage of abortive fruit. Cauline leaves are much reduced and arise from the base of the stem, giving an almost acaulescent appearance to the plant. Breeding studies (J. G. Harris 1985) indicate that subspp. maccallae and porsildii are self-incompatible, while most subspecies of Braya humilis are strongly autogamous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 17 (7 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Plants not scapose; cauline leaves (1 or) 2-4; fruits linear; seeds usually uniseriate, rarely weakly biseriate
→ 2
1. Plants scapose; cauline leaves 0 or 1 (or with leaflike bract subtending proximalmost pedicel); fruits ovoid, globose, oval-elliptic, oblong, oblong-elliptic, oblong-lanceoloid, or lanceoloid-subulate, not linear; seeds usually biseriate (uniseriate in B. fernaldii and, sometimes, B. longii)
→ 3
2. Cauline leaves 3 or more; basal leaves (0.3-)0.5-2(-3.5) cm × 1-8(-10) mm; racemes elongated in fruit; fruits (0.9-)1.2-2.5(-3.2) cm
B. humilis
2. Cauline leaves 1-4; basal leaves 0.5-3 cm × 0.5-2(-3) mm; racemes not elongated in fruit; fruits (0.5-)0.9-1.2(-1.4) cm.
B. linearis
3. Fruits ovoid or globose
→ 4
3. Fruits oval-elliptic, oblong, oblong-elliptic, oblong-lanceoloid, or lanceoloid-subulate
→ 5
4. Petals 4.7-6.6 × 3-5.1 mm; styles 1.2-2(-2.5) mm; stems erect to ascending.
B. pilosa
4. Petals 2-3.7 × 1-1.5 mm; styles obsolete to 0.7(-1) mm; stems usually decumbent to prostrate, sometimes ascending.
B. thorild-wulffii
5. Fruits oval-elliptic, oblong-cylindrical, or lanceoloid; septum margins not expanded basally (not forming sacklike pouch around proximalmost seeds); seeds biseriate.
B. glabella
5. Fruits lanceoloid-subulate; septum margins broadly expanded basally (forming sacklike pouch around proximalmost seeds); seeds somewhat to nearly uniseriate
→ 6
6. Fruit valves pubescent; petals 2.4-3.8(-4) × (0.8-)1-1.3(-2) mm, (claws often not well- differentiated from blades).
B. fernaldii
6. Fruit valves glabrous or sparsely pubescent; petals (3-)3.3-4.8(-5) × (1.2-)1.4-2.5(-3) mm, (claws usually well-differentiated from blades).
B. longii
Source FNA vol. 7, p. 550. FNA vol. 7, p. 546. Author: James G. Harris.
Parent taxa Brassicaceae > tribe Euclidieae > Braya > Braya humilis Brassicaceae > tribe Euclidieae
Sibling taxa
B. humilis subsp. ellesmerensis, B. humilis subsp. humilis, B. humilis subsp. porsildii
Subordinate taxa
B. fernaldii, B. glabella, B. humilis, B. linearis, B. longii, B. pilosa, B. thorild-wulffii
Synonyms Platypetalum
Name authority J. G. Harris: Novon 16: 346. (2006) Sternberg & Hoppe: Denkschr. Königl.-Baier. Bot. Ges. Regensburg 1(1): 65. (1815)
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