Braya humilis subsp. maccallae |
Braya |
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dwarf braya, maccall's dwarf braya, mccalla's dwarf braya |
northern-rockcress |
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Habit | Perennials; (sometimes pulvinate, caudex simple or many-branched); scapose or not; usually pubescent or pilose, sometimes glabrous, trichomes short-stalked, forked, subdendritic, or submalpighiaceous, mixed with simple ones (rarely exclusively). | |||||||||||||||||||||||||
Stems | ascending to erect, often unbranched, 0.4–2.3 dm, moderately pubescent. |
erect to decumbent or ascending, unbranched or branched. |
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Leaves | blade margins entire or, sometimes, sinuate-dentate, surfaces glabrescent to moderately pubescent. |
basal and, sometimes, cauline; petiolate or sessile; basal rosulate, petiolate, blade margins entire, sinuate, dentate, or, rarely, pinnately lobed; cauline usually absent, rarely few present, (sub)sessile, blade margins usually entire, rarely dentate or pinnately lobed. |
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Racemes | (corymbose, sometimes bracteate basally or throughout), elongated or not in fruit. |
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Flowers | petals white, (4.4–)4.9–6.7(–7.2) × (2.1–)2.3–4(–4.2) mm. |
sepals [sometimes persistent], oblong [ovate], lateral pair not saccate basally (sometimes slightly so in B. humilis and B. linearis); petals white, pink, or purple [rarely pale yellow], obovate, oblanceolate, or spatulate, (slightly to much longer than sepals), claw distinct or not, (shorter than sepal, apex obtuse or rounded); stamens tetradynamous; filaments dilated or not basally; anthers ovate or oblong, (apex usually obtuse, sometimes apiculate); nectar glands (4), lateral, 1 on each side of lateral stamen. |
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Fruiting pedicels | erect, divaricate, or ascending, slender (much narrower than fruit). |
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Fruits | often abortive, somewhat torulose, 0.7–1.2 mm wide when fully developed; septum not fenestrate or split longitudinally. |
siliques or silicles, sessile, linear, oblong, cylindrical, oval-elliptic, ovoid, lanceoloid, lanceoloid-subulate, or globose, smooth or torulose, terete or slightly latiseptate; valves each often with prominent midvein, glabrous or pubescent; replum rounded; septum complete, (membranous, translucent); ovules (5–)14–44 per ovary; stigma capitate, entire or slightly 2-lobed. |
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Seeds | plump, not winged, oblong or ovoid; seed coat (minutely reticulate), not mucilaginous when wetted; cotyledons incumbent. |
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x | = 7. |
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2n | = 28. |
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Braya humilis subsp. maccallae |
Braya |
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Phenology | Flowering May–Jul. | |||||||||||||||||||||||||
Habitat | Sandy gravelly riverbanks and floodplains, sometimes on slopes and glacial moraines | |||||||||||||||||||||||||
Elevation | 1500-3000 m (4900-9800 ft) | |||||||||||||||||||||||||
Distribution |
AB; BC |
North America; n Europe; Asia |
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Discussion | Subspecies maccallae differs from subspp. elesmerensis and humilis in having leaf margins mostly entire, flowers exceptionally large, and a high percentage of abortive fruit. Cauline leaves are much reduced and arise from the base of the stem, giving an almost acaulescent appearance to the plant. Breeding studies (J. G. Harris 1985) indicate that subspp. maccallae and porsildii are self-incompatible, while most subspecies of Braya humilis are strongly autogamous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species 17 (7 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 7, p. 550. | FNA vol. 7, p. 546. | ||||||||||||||||||||||||
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Synonyms | Platypetalum | |||||||||||||||||||||||||
Name authority | J. G. Harris: Novon 16: 346. (2006) | Sternberg & Hoppe: Denkschr. Königl.-Baier. Bot. Ges. Regensburg 1(1): 65. (1815) | ||||||||||||||||||||||||
Web links |