Braya humilis subsp. humilis |
Brassicaceae tribe Euclidieae |
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dwarf braya |
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Habit | Annuals or perennials [shrubs or subshrubs]; eglandular. | |
Stems | ascending to erect, branched or unbranched, 0.4–3.3 dm, sparsely to moderately pubescent. |
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Leaves | blade margins sinuate-dentate, shallowly pinnatifid, or entire, surfaces moderately pubescent. |
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Cauline leaves | (sometimes absent), usually petiolate or sessile, sometimes subsessile; blade base not auriculate, margins usually entire or dentate, rarely lobed. |
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Trichomes | short-stalked or sessile, stellate, forked, dendritic, malpighiaceous, or simple, rarely absent. |
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Racemes | usually ebracteate, usually elongated in fruit. |
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Flowers | petals white, pink, or purple, 2.5–6.9(–7.5) × (0.7–)0.9–4(–4.2) mm. |
actinomorphic; sepals erect [ascending to spreading], lateral pair seldom saccate basally; petals white, pink, or purple [yellow], claw present, distinct or obscure; filaments unappendaged, not winged; pollen 3-colpate. |
Fruits | usually fertile and fully developed, somewhat torulose, 0.6–1.2(–1.3) mm wide; septum not fenestrate or split longitudinally. |
silicles or siliques, usually dehiscent, unsegmented, terete or latiseptate [angustiseptate]; ovules 2–80[–numerous] per ovary; style distinct or obsolete; stigma entire or strongly 2-lobed. |
Seeds | usually biseriate or uniseriate (aseriate in Euclidium); cotyledons accumbent or incumbent. |
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2n | = 28, 42, 56, 70. |
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Braya humilis subsp. humilis |
Brassicaceae tribe Euclidieae |
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Phenology | Flowering May–Jul. | |
Habitat | Sandy, gravelly soil along streams, lakeshores, roadsides, moraines, open stony slopes, dolomite cliffs and slopes, limestone ledges, solifluction soils | |
Elevation | 0-4000 m (0-13100 ft) | |
Distribution |
AK; CO; MI; MT; VT; WY; AB; BC; MB; NF; NT; NU; ON; QC; YT; Greenland; e Asia; c Asia |
North America; Europe; Asia; n Africa |
Discussion | Subspecies humilis is extremely variable morphologically. In a general way, morphological form correlates with ploidy level, e.g., tetraploids, octoploids, and decaploids tend to be short in stature with small leaves. Hexaploids are less predictable. They range from short plants with small leaves to large, robust, multi-branched plants with large, pinnatifid leaves. Attempting to segregate most morphological forms of Braya humilis into logical infraspecific taxa is an exercise in futility. Populations that appear distinctive in the field almost always blur imperceptibly into the larger subsp. humilis continuum when compared with other populations from across the range of distribution. Subspecies humilis is broadly distributed on calcareous substrates in arctic, subarctic, alpine, and boreal regions of North America and Asia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Genera 13, species 115 (3 genera, 9 species in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 7, p. 549. | FNA vol. 7, p. 545. |
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Synonyms | Arabidopsis novae-angliae, B. humilis var. abbei, B. humilis subsp. arctica, B. humilis var. arctica, B. humilis var. interior, B. humilis var. laurentiana, B. humilis var. leiocarpa, B. humilis var. novae-angliae, B. humilis subsp. richardsonii, B. humilis subsp. ventosa, B. humilis var. ventosa, B. intermedia, B. novae-angliae, B. novae-angliae subsp. abbei, B. novae-angliae var. interior, B. novae-angliae var. laurentiana, B. novae-angliae subsp. ventosa, B. richardsonii, Pilosella novae-angliae, Pilosella richardsonii, Torularia humilis subsp. arctica | |
Name authority | unknown | de Candolle: Mém. Mus. Hist. Nat. 7: 236. (1821) |
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