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alpine braya, smooth northern-rockcress

Photo is of parent taxon

smooth braya, smooth northern-rockcress

Habit Plants scapose; sparsely to densely pubescent, trichomes simple, 2- or 3-forked.
Stems

simple or few to several from base, ascending or erect, rarely decumbent to prostrate, (0.1–)0.3–1.7(–2.3) dm.

ascending to erect, 0.5–1.7(–2.3) dm.

Leaves

blade often somewhat fleshy, to 4 mm wide, margins often entire, sometimes 1 or 2 weak teeth per side.

Basal leaves

blade (often somewhat fleshy), linear-oblanceolate to broadly spatulate, (0.4–)0.8–6(–7.9) cm × (0.3–)0.6–4(–6) mm, base (membranous), broadly expanded near point of attachment, margins usually entire, sometimes weakly dentate with 1 or 2 teeth per side, apex obtuse, (often with a tuft of long, simple hairs, surfaces sparsely to moderately pubescent).

Cauline leaves

0 or 1 (or a leaflike bract subtending proximalmost pedicel). (Racemes elongated or not in fruit.) Fruiting pedicels ascending-erect to divaricate, (0.9–)1.9–7.5(–8.6) mm.

Racemes

often loosely elongated in fruit.

Flowers

sepals (1.6–)1.9–3.7 × (0.7–)1–2 mm;

petals white or purplish (broadly obovate or spatulate), (2.1–)2.4–4.5(–4.7) × (0.7–)1–3(–3.2) mm, (apex rounded);

filaments 1.7–2.2 mm;

anthers oblong, 0.3–0.5 mm.

Fruits

oval-elliptic, oblong-elliptic, oblong, or oblong-lanceoloid, sometimes slightly torulose, (straight or somewhat curved), (0.3–)0.5–1.2(–1.5) cm × (0.8–)1.1–3(–3.6) mm;

valves sparsely to densely pubescent or glabrous, trichomes simple or 2 (or 3)-forked;

septum margin not expanded or not basally;

ovules (5–)16–20 per ovary;

style (0.3–)0.5–1.8(–2) mm;

stigma entire or strongly 2-lobed, (narrow or broad).

oblong to narrowly oblong-lanceoloid, often curved, (0.3–)0.5–1.2(–1.5) cm × (0.8–)1.1–3(–3.6) mm, 3.5–8.3 times as long as wide;

style slender or stout, (0.3–)0.5–1.6(–2) mm.

Seeds

biseriate, oblong, (0.7–)0.9–1.6 × 0.4–0.8(–0.9) mm.

2n

= 56.

Braya glabella

Braya glabella subsp. glabella

Phenology Flowering Jun–Jul.
Habitat Barren, usually calcareous soils and gravel, gravel bars, disturbed sites, lake and sea shores, scree slopes, solifluction lobes
Elevation 0-3700 m (0-12100 ft)
Distribution
from FNA
AK; CO; MT; WY; AB; BC; NT; NU; QC; YT; Asia
[BONAP county map]
from FNA
AK; CO; MT; WY; AB; BC; NT; NU; QC; YT; Asia
[BONAP county map]
Discussion

Subspecies 3 (3 in the flora).

Braya glabella is extremely variable, sometimes even within a population, and the species has often been split into taxa based primarily on fruit attributes. When the species is examined from its entire range, the perceived morphological gaps blur into a bewildering array of overlapping forms. On the basis of morphological and molecular evidence (S. I. Warwick et al. 2004) it is difficult to justify the recognition of more than one species in this very plastic group, but it does seem useful to divide the species into three fairly distinctive subspecies. Some populations will be readily separable into one of these subspecies, but others will likely defy unequivocal placement, particularly those from areas where the ranges of the subspecies meet.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruits oblong to narrowly oblong-lanceoloid, 3.5-8.3 times as long as wide; racemes often loosely elongated in fruit.
subsp. glabella
1. Fruits oval-elliptic to oblong-elliptic, rarely broadly oblong-lanceoloid, 2.5-3.7 times as long as wide; racemes not elongated in fruit, often compact
→ 2
2. Stems decumbent to prostrate, sometimes weakly ascending; leaf blades not fleshy, to 6 mm wide; fruits 0.8-1.2 cm; styles 0.8-1.8 mm.
subsp. prostrata
2. Stems ascending to erect; leaf blades often fleshy, to 4 mm wide; fruits 0.5-1 cm; styles 0.5- 1.2 mm.
subsp. purpurascens
Source FNA vol. 7, p. 547. FNA vol. 7, p. 548.
Parent taxa Brassicaceae > tribe Euclidieae > Braya Brassicaceae > tribe Euclidieae > Braya > Braya glabella
Sibling taxa
B. fernaldii, B. humilis, B. linearis, B. longii, B. pilosa, B. thorild-wulffii
B. glabella subsp. prostrata, B. glabella subsp. purpurascens
Subordinate taxa
B. glabella subsp. glabella, B. glabella subsp. prostrata, B. glabella subsp. purpurascens
Synonyms B. alpina var. glabella B. alpina var. americana, B. americana, B. bartlettiana, B. bartlettiana var. vestita, B. henryae, B. humilis var. americana
Name authority Richardson: in J. Franklin et al., Narr. Journey Polar Sea, 743. (1823) unknown
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