Bellardia |
Bellardia latifolia |
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bellardia, glandweed, lineseed, parentucellia |
broadleaf glandweed or parentucellia |
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Habit | Herbs, annual [perennial]; hemiparasitic. | |||||||||
Stems | erect, not fleshy, retrorsely short-strigose or glandular-hairy. |
simple or with few ascending branches proximally, 6–30 cm, glandular-hairy. |
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Leaves | cauline, opposite, sometimes some subopposite or alternate; petiole absent; blade not fleshy, not leathery, margins coarsely crenate-dentate. |
4 or 5 pairs, ascending or divaricate, glandular-hairy; blade ovate, 4–10(–12) x 2–7 mm, margins purple, apex acute. |
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Inflorescences | terminal, spikelike racemes; bracts present. |
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Pedicels | present; bracteoles absent. |
0.5–2 mm, glabrous. |
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Flowers | sepals 4, calyx radially or bilaterally symmetric, not flattened laterally, tubular or campanulate, not accrescent in fruit, lobes triangular or lanceolate; petals 5, corolla white with purple galea, yellow, or red-purple, strongly bilabiate, tubular-funnelform, abaxial lobes 3, adaxial 2, adaxial lip galeate; stamens 4, didynamous, filaments glabrous, anther mucros equal or absent; staminode 0; ovary 2-locular, placentation axile; stigma clavate, capitate, or +/- 2-lobed. |
calyx tubular, 6–12 mm, tube 4–9.5 mm, hairy or glandular-hairy, lobes +/- equal, lanceolate, 2–4 x 1.5–2 mm, herbaceous, margins entire, apex acute, hairy or glandular-hairy; corolla red-purple, 8–18 mm, glandular-hairy externally, throat with 2 yellow, ridgelike appendages, abaxial lobes spreading, adaxial projecting; stamens included, pollen sacs yellow, 1–2 mm, mucronate distally, glabrous, dehiscing longitudinally in distal 3/4–4/5; style 4–6 mm, glabrous; stigma 2-lobed. |
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Capsules | dehiscence loculicidal. |
6–12 x 2–3 mm, glabrous. |
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Seeds | 150–450, white, reddish brown with age, ellipsoid or ellipsoid-oblong, wings absent. |
0.3–0.5 mm, smooth. |
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Spikelike | racemes 2–18 cm; flowers 1–12 pairs, interrupted proximally, dense distally, glandular-hairy; peduncle absent; bracts foliaceous, 7–9 x 6–7 mm, margins of distal bracts with 1 pair of teeth. |
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x | = 12. |
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2n | = 48 (Europe). |
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Bellardia |
Bellardia latifolia |
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Phenology | Flowering Apr–May. | |||||||||
Habitat | Pastures, roadsides, open areas. | |||||||||
Elevation | 0–100 m. (0–300 ft.) | |||||||||
Distribution |
Europe [Introduced in North America; introduced also in s South America, Asia, Africa, Atlantic Islands, Pacific Islands, s Australia] |
CA; Europe; n Africa; Atlantic Islands (Canary Islands) [Introduced in North America; introduced also in South America, w Asia, s Australia] |
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Discussion | Species 48 (3 in the flora). Bellardia trixago has been included in a monospecific section of Bartsia in the broad sense (U. Molau 1990). Morphology and molecular phylogenetic analyses indicate a close relationship among Bellardia, Parentucellia, and New World species of Bartsia (A. D. Wolfe et al. 2005; J. R. Bennett and S. Mathews 2006; S. Uribe-Convers and D. C. Tank, http://2010.botanyconference.org/engine/search/index.php?func=detail&aid=685; A. Scheunert et al. 2012). Molecular phylogenetic analyses suggest that South American Bartsia are nested within a highly supported clade including Bellardia and Parentucellia (Scheunert et al.). An expanded Bellardia includes P. latifolia (Linnaeus) Caruel, P. viscosa (Linnaeus) Caruel, and the South American species of Bartsia. After this treatment had been completed, the author became aware of the alternative view presented by Uribe-Convers and Tank (2016), where all of the South American Bartsia taxa are transferred to the new genus Neobartsia Uribe-Convers & Tank, Parentucellia is recognized, and Bellardia includes only B. trixago and B. viscosa. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Bellardia latifolia may be found also in the Canary Islands and as an introduction to South America. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 17, p. 490. | FNA vol. 17, p. 490. | ||||||||
Parent taxa | ||||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Synonyms | Bartsia section bellardia, Parentucellia | Euphrasia latifolia, Parentucellia latifolia | ||||||||
Name authority | Allioni: Fl. Pedem. 1: 61. (1785) | (Linnaeus) Cuatrecasas: Trab. Mus. Ci. Nat. Barcelona 12: 428. (1929) | ||||||||
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