Flora of North America species comparison

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Bellardia

bellardia, glandweed, lineseed, parentucellia
Habit Herbs, annual [perennial]; hemiparasitic.
Stems

erect, not fleshy, retrorsely short-strigose or glandular-hairy.
Leaves

cauline, opposite, sometimes some subopposite or alternate;

petiole absent;

blade not fleshy, not leathery, margins coarsely crenate-dentate.
Inflorescences

terminal, spikelike racemes;

bracts present.
Pedicels

present;

bracteoles absent.
Flowers

sepals 4, calyx radially or bilaterally symmetric, not flattened laterally, tubular or campanulate, not accrescent in fruit, lobes triangular or lanceolate;

petals 5, corolla white with purple galea, yellow, or red-purple, strongly bilabiate, tubular-funnelform, abaxial lobes 3, adaxial 2, adaxial lip galeate;

stamens 4, didynamous, filaments glabrous, anther mucros equal or absent;

staminode 0;

ovary 2-locular, placentation axile;

stigma clavate, capitate, or +/- 2-lobed.
Capsules

dehiscence loculicidal.
Seeds

150–450, white, reddish brown with age, ellipsoid or ellipsoid-oblong, wings absent.
x

= 12.

Bellardia
Distribution
from USDA
Europe [Introduced in North America; introduced also in s South America, Asia, Africa, Atlantic Islands, Pacific Islands, s Australia]
[BONAP county map]
Discussion

Species 48 (3 in the flora).

Bellardia trixago has been included in a monospecific section of Bartsia in the broad sense (U. Molau 1990). Morphology and molecular phylogenetic analyses indicate a close relationship among Bellardia, Parentucellia, and New World species of Bartsia (A. D. Wolfe et al. 2005; J. R. Bennett and S. Mathews 2006; S. Uribe-Convers and D. C. Tank, http://2010.botanyconference.org/engine/search/index.php?func=detail&aid=685; A. Scheunert et al. 2012). Molecular phylogenetic analyses suggest that South American Bartsia are nested within a highly supported clade including Bellardia and Parentucellia (Scheunert et al.). An expanded Bellardia includes P. latifolia (Linnaeus) Caruel, P. viscosa (Linnaeus) Caruel, and the South American species of Bartsia. After this treatment had been completed, the author became aware of the alternative view presented by Uribe-Convers and Tank (2016), where all of the South American Bartsia taxa are transferred to the new genus Neobartsia Uribe-Convers & Tank, Parentucellia is recognized, and Bellardia includes only B. trixago and B. viscosa.

(Discussion copyrighted by Flora of North America; reprinted with permission.)
Key
1. Corollas white with purple galea; calyx lobes unequal.
B. trixago
1. Corollas red-purple or yellow; calyx lobes +/- equal.
→ 2
2. Corollas red-purple; leaf blades 4–10(–12) mm.
B. latifolia
2. Corollas yellow; leaf blades (10–)20–46 mm.
B. viscosa
Source FNA vol. 17, p. 490. Author: Elizabeth H. Zacharias.
Parent taxa Orobanchaceae
Subordinate taxa
B. latifolia, B. trixago, B. viscosa
Synonyms Bartsia section bellardia, Parentucellia
Name authority Allioni: Fl. Pedem. 1: 61. (1785)
Web links 
Unless otherwise noted, all data comes from the online version of the Flora of North America, who have made most of their content available via a Creative Commons license (CC BY 4.0).



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