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broadscale, mound saltbush, New Mexico saltbush, silver saltbush

four-wing saltbu sh, four-wing shad-scale, hoary saltbush, shadscale, wingscale

Habit Subshrubs, dioecious, clump forming, mainly 2–8 dm and as wide, woody at base. Shrubs, dioecious or rarely monoecious, mainly 8–20 dm, as wide or wider, not especially armed.
Stems

stiffly erect;

branchlets terete.

Leaves

tardily deciduous, alternate or proximal-most subopposite, shortly petiolate;

blade gray green, oblong-ovate to elliptic or orbiculate, 8–30(–35) × 6–20 mm, margin entire or rarely dentate, apex rounded to retuse or obtuse.

persistent, alternate, sessile or nearly so, blade linear to oblanceolate, oblong, or obovate, mainly 10–40 × 3–8 mm, margin entire, apex retuse to obtuse.

Staminate flowers

yellow, in clusters 2–3 mm wide, borne in panicles 6–30 cm.

yellow (rarely brown), in clusters 2–3 mm wide, borne in panicles 3–15 cm.

Pistillate flowers

in small, very numerous glomerules in axils of elongated, terminal leafy-bracteate spikes or finally paniculate.

borne in panicles 5–40 cm.

Seeds

brown, 2.4–2.8 mm.

1.5–2.5 mm wide.

Fruiting

bracteoles sessile or substipitate, 4–5 × 5–9 mm, base broadly cuneate, margin sharply toothed, apical tooth subtended by 2–6 equal or smaller teeth, faces smooth or rarely tuberculate.

bracteoles 8–25 mm, as wide, on stipes 1–8 mm, with 4 prominent wings extending the bract length, united throughout, wings dentate to entire, apex toothed, surface of wings and body smooth or reticulate.

2n

= 18, 36+.

Atriplex obovata

Atriplex canescens

Phenology Flowering summer–fall.
Habitat Fine-textured substrates, with salt desert shrub and lower pinyon-juniper communities
Elevation 1500-2000 m (4900-6600 ft)
Distribution
from FNA
AZ; CO; NM; TX; UT; Mexico
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CO; ID; KS; MT; ND; NE; NM; NV; OK; OR; SD; TX; UT; WA; WY; AB; Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

This species forms hybrids with Atriplex confertifolia and A. gardneri varieties (see var. bonnevillensis). Materials from the vicinity of the type locality of the species in South Dakota are low subherbaceous plants that differ from our shrubby tall material. However, the type area is presently covered with water from a dam on the Missouri River, and it is not possible to exclude the possibility of A. canescens as it has been interpreted for the past century to have existed at that site during the Lewis and Clark Expedition, if that is indeed where the lectotype was collected.

Varieties 4 (4 in the flora).

Nuttall’s new combination Atriplex canescens was based on Calligonum canescens Pursh. Watson based his new name A. nuttallii directly on A. canescens Nuttall, i.e., including the citation of Nuttall’s “Genera, 1. 197,” and on its basionym, C. canescens. The name A. nuttallii is thus a nomenclatural synonym of A. canescens and was thus illegitimate at its inception. It cannot be resurrected by even the most sophisticated arguments. A sheet of Atriplex canescens, noted as “a shrub,” taken by Nuttall on the 1810 Missouri River expedition is extant in the Lambert herbarium (PH). It bears several, obviously shrubby, staminate flowering branches, but the only pistillate branch is very immature. The name A. gardneri, also cited provisionally by Watson within the concept of A. nuttallii, clearly has priority over other names for that widely distributed species complex. Attempts at leptotypification of the name nuttallii by J. McNeill et al. (1983) and H. C. Stutz and S. C. Sanderson (1998) are both superfluous, the name being illegitimate.

C. A. Hanson (1962) noted the similarity between the occasional wingless fruiting bracteoles of Atriplex canescens and A. gardneri var. falcata. He noted further that the bracts of both species lack lateral teeth subtending the terminal ones, have terminal teeth united half their length, and have indurate bracts. Whether such similarity indicates relationship or mere coincidence is open to question. However, A. canescens is known to form hybrids with most, if not all, portions of the gardneri complex and with other woody species whose range it overlaps as well.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Fruiting bracteoles merely dentate to entire, commonly 8-25 mm wide; plants of broad or other distribution
→ 2
1. Fruiting bracteoles dentate to laciniate, mainly 5-10 mm wide; plants of s California
→ 3
2. Bracteoles mainly less than 12 mm wide; plants not layering in sand dunes, widespread
var. canescens
2. Bracteoles mainly 15- mm wide; plants layering in sand dunes, cw Utah
var. gigantea
3. Fruiting bracteoles mainly 4-8 mm wide, dentate or laciniate; shrubs mainly 3-10 dm
var. macilenta
3. Fruiting bracteoles mainly 8-11 mm wide, la- ciniate; shrubs mainly 10-20 dm
var. laciniata
Source FNA vol. 4, p. 371. FNA vol. 4.
Parent taxa Chenopodiaceae > Atriplex > subg. Pterochiton Chenopodiaceae > Atriplex > subg. Pterochiton
Sibling taxa
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. canescens, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
A. acanthocarpa, A. amnicola, A. argentea, A. californica, A. confertifolia, A. cordulata, A. coronata, A. corrugata, A. coulteri, A. covillei, A. dioica, A. elegans, A. fruticulosa, A. gardneri, A. garrettii, A. glabriuscula, A. gmelinii, A. graciliflora, A. heterosperma, A. holocarpa, A. hortensis, A. hymenelytra, A. joaquiniana, A. klebergorum, A. laciniata, A. lentiformis, A. leucophylla, A. lindleyi, A. linearis, A. littoralis, A. matamorensis, A. mucronata, A. nudicaulis, A. nummularia, A. oblongifolia, A. obovata, A. pacifica, A. parishii, A. parryi, A. patula, A. pentandra, A. phyllostegia, A. pleiantha, A. polycarpa, A. powellii, A. prostrata, A. pusilla, A. rosea, A. saccaria, A. semibaccata, A. serenana, A. spinifera, A. suberecta, A. suckleyi, A. tatarica, A. torreyi, A. truncata, A. tularensis, A. watsonii, A. wolfii, A. wrightii
Subordinate taxa
A. canescens var. canescens, A. canescens var. gigantea, A. canescens var. laciniata, A. canescens var. macilenta
Synonyms A. greggii, A. jonesii, A. obovata var. tuberata Calligonum canescens, A. nuttallii
Name authority Moquin-Tandon: Chenop. Monogr. Enum., 61. (1840) (Pursh) Nuttall: Gen. N. Amer. Pl. 1: 197. (1818)
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