Burke: Cardamine nuttallii vs. Cardamine angulata

	Cardamine nuttallii	Cardamine angulata
	beautiful bittercres, Nuttall's toothwort, slender toothwort	angled bittercress, seaside bittercress
Habit	Glabrous to sparsely pubescent perennial from short, slender, fleshy rhizomes, the stems erect, 1-2 dm. tall.	Perennial from spreading rhizomes, the simple stem 2-8 dm. tall, with stiff, simple hairs at least at the base.
Leaves	Basal leaves long-petiolate, the blades orbicular to cordate, usually simple and scalloped, if deeply lobed the lobes entire; cauline leaves 1-3, grouped above mid-length on the stem, usually with 3-5 lanceolate, entire leaflets 1-4 cm. long.	Leaves mainly cauline, alternate, trifoliate, long-petiolate, the leaflets ovate to ovate-lanceolate, 1.5-7 cm. long, 3- to 5-lobed or toothed, short-petiolate, the terminal leaflet the largest.
Flowers	Inflorescence of bractless, few-flowered racemes; pedicles ascending to erect, 1-2 cm. long; sepals 4, 3-5 mm. long, purplish, the outer pair saccate at the base; petals 4, pink to reddish or purplish, 9-14 mm. long; stamens 6; style slender, 3-6 mm. long.	Flowers in bractless racemes; pedicles ascending, 1-2 cm. long; sepals 4, oval, 2.5-3 mm. long, the outer pair saccate at the base; petals 4, white to pinkish, 8-13 mm. long, somewhat clawed; stamens 6; style 1.5-4 mm. long.
Fruits	Siliques linear, 1.5-5 cm. long and 1.5 mm. broad; seeds in 1 series.	Siliques linear, erect, 2-4 cm. long and 2 mm. broad; seed in1 series.

	Cardamine nuttallii	Cardamine angulata
Flowering time	March-May	April-June
Habitat	Seasonally moist soils of forest openings to forest understory, from low to moderate elevations.	Wet ground, especially along stream banks, often in deep woods.
Distribution	Occurring on both sides of the Cascades crest in Washington; British Columbia to California. [WildflowerSearch map] [BONAP county map]	Occurring west of the Cascades crest in Washington; Alaska to northern California. [WildflowerSearch map] [BONAP county map]
Origin	Native	Native
Conservation status	Not of concern	Not of concern
Sibling taxa	C. angulata, C. bellidifolia, C. breweri, C. cordifolia, C. corymbosa, C. flexuosa, C. hirsuta, C. occidentalis, C. occulta, C. oligosperma, C. pensylvanica, C. pratensis, C. umbellata	C. bellidifolia, C. breweri, C. cordifolia, C. corymbosa, C. flexuosa, C. hirsuta, C. nuttallii, C. occidentalis, C. occulta, C. oligosperma, C. pensylvanica, C. pratensis, C. umbellata
Web links	Local floras: BC, CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search	Local floras: BC, CA, OR, WA Local Web sites: CalFlora, CalPhotos, Flora NW, PNW Herbaria, Turner Photog. WildflowerSearch iNaturalist (observations) USDA Plants Database LBJ Wildflower Center SEINet Plants of the World Online Encyclopedia of Life Wikipedia Google Image Search

Cardamine nuttallii

Cardamine angulata

beautiful bittercres, Nuttall's toothwort, slender toothwort

angled bittercress, seaside bittercress

Habit

Glabrous to sparsely pubescent perennial from short, slender, fleshy rhizomes, the stems erect, 1-2 dm. tall.

Perennial from spreading rhizomes, the simple stem 2-8 dm. tall, with stiff, simple hairs at least at the base.

Leaves

Basal leaves long-petiolate, the blades orbicular to cordate, usually simple and scalloped, if deeply lobed the lobes entire;

cauline leaves 1-3, grouped above mid-length on the stem, usually with 3-5 lanceolate, entire leaflets 1-4 cm. long.

Leaves mainly cauline, alternate, trifoliate, long-petiolate, the leaflets ovate to ovate-lanceolate, 1.5-7 cm. long, 3- to 5-lobed or toothed, short-petiolate, the terminal leaflet the largest.

Flowers

Inflorescence of bractless, few-flowered racemes; pedicles ascending to erect, 1-2 cm. long;

sepals 4, 3-5 mm. long, purplish, the outer pair saccate at the base;

petals 4, pink to reddish or purplish, 9-14 mm. long;

stamens 6;

style slender, 3-6 mm. long.

Flowers in bractless racemes; pedicles ascending, 1-2 cm. long;

sepals 4, oval, 2.5-3 mm. long, the outer pair saccate at the base;

petals 4, white to pinkish, 8-13 mm. long, somewhat clawed;

stamens 6;

style 1.5-4 mm. long.

Fruits