Cardamine nuttallii |
Cardamine angulata |
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Nuttall's toothwort, slender toothwort |
angle-leaved bittercress, angled bittercress, seaside bittercress |
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Habit | Plants perennial, glabrous or pubescent; rhizomes slender, tuberous, segmented, producing fragile segments, ovoid, oblong, or cylindrical, 2–5 mm in diameter, nodes fleshy. | Plants perennial, glabrous or pubescent; rhizomes slender, cylindrical, to 2 mm in diameter. |
Stems | erect, 0.5–2(3) dm; simple, bases above ground much wider than attachment to rhizome. |
(15)25–85(100) cm, bases above ground as wide as its attachment to rhizome; hirsute basally. |
Rhizomal leaves | simple or 3(5)-foliolate; (3)4–20(25) cm; blades or terminal leaflets reniform, suborbicular; ovate, or oblong; (0.9)1.3–4(5.2) × (0.8)1.2–5(7) cm, often with purple spots, bases cordate to obtuse; margins apiculate-dentate or 5–7-lobed. |
3(5)-foliolate; (4)7–20(22) cm, bases cuneate, rarely subreniform or obtuse; margins 3–5(7)-lobed or -toothed, puberulent; terminal leaflets ovate to lanceolate, 1.5–7(9) cm. |
Cauline leaves | 1–3, pinnately or palmately compound, 3(5)-foliolate, bases not auriculate; terminal leaflets ovate, oblong, or linear; (0.5)1–3.5(6) cm; lateral leaflets smaller; entire, dentate, or lobed. |
(3)4–8, pinnately or palmately compound, 3(5)-foliolate, bases not auriculate, pubescent along margins; terminal leaflets ovate to lanceolate, 2–7 × 0.6–4 cm. |
Inflorescences | bracts 0, fruiting pedicels ascending to divaricate, 1–3.5 cm. |
bracts 0, fruiting pedicels ascending to divaricate; (9)12–25 mm. |
Flowers | sepals erect, oblong, 3.5–5 mm; lateral pair saccate; petals obovate, 10–15 × 4–7.5 mm, purple to pale pink, rarely white, not clawed; tips rounded; ovules 8–16 per ovary; styles 4–8 mm. |
sepals oblong, 2.5–4 mm; lateral pair saccate; petals white or rarely pinkish, obovate, 8–15 × 4–8 mm, clawed; tips rounded or emarginate; ovules 10–16 per ovary; styles (0.5)1–4 mm. |
Fruits | 25–56 × 2–2.3 mm, glabrous. |
15–32 × 1.4–2 mm, glabrous. |
Seeds | oblong, 2–2.5 × 1.4–1.6 mm. |
oblong, 1.8–2.3 × 1–1.2 mm. |
2n | =40. |
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Cardamine nuttallii |
Cardamine angulata |
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Distribution | ||
Discussion | Damp woods, shaded bottomlands and hillsides, mossy slopes, streamsides. Flowering Mar–Jul. 0–2000 m. Casc, Col, CR, ECas, Est, Sisk, WV. CA, WA; north to British Columbia. Native. |
Wet meadows, moist grounds, thickets, streambanks, swampy or damp woods. Flowering Apr–Jun. 0–1400 m. Casc, CR, Est, Sisk, WV. CA, WA; north to AK. Native. |
Source | Flora of Oregon, volume 2 draft Ihsan Al-Shehbaz |
Flora of Oregon, volume 2, page 464 Ihsan Al-Shehbaz |
Sibling taxa | ||
Synonyms | Cardamine nuttallii var. covilleana, Cardamine nuttallii var. dissecta, Cardamine nuttallii var. gemmata, Cardamine nuttallii var. nuttallii, Cardamine pulcherrima, Cardamine pulcherrima var. pulcherrima, Cardamine pulcherrima var. tenella, Dentaria gemmata, Dentaria tenella, Dentaria tenella var. pulcherrima, Dentaria tenella var. quercetorum, Dentaria tenella var. tenella | |
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