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Nuttall's toothwort, slender toothwort

angle-leaved bittercress, angled bittercress, seaside bittercress

Habit Plants perennial, glabrous or pubescent; rhizomes slender, tuberous, segmented, producing fragile segments, ovoid, oblong, or cylindrical, 2–5 mm in diameter, nodes fleshy. Plants perennial, glabrous or pubescent; rhizomes slender, cylindrical, to 2 mm in diameter.
Stems

erect, 0.5–2(3) dm; simple, bases above ground much wider than attachment to rhizome.

(15)25–85(100) cm, bases above ground as wide as its attachment to rhizome; hirsute basally.

Rhizomal leaves

simple or 3(5)-foliolate; (3)4–20(25) cm;

blades or terminal leaflets reniform, suborbicular; ovate, or oblong; (0.9)1.3–4(5.2) × (0.8)1.2–5(7) cm, often with purple spots, bases cordate to obtuse;

margins apiculate-dentate or 5–7-lobed.

3(5)-foliolate; (4)7–20(22) cm, bases cuneate, rarely subreniform or obtuse;

margins 3–5(7)-lobed or -toothed, puberulent;

terminal leaflets ovate to lanceolate, 1.5–7(9) cm.

Cauline leaves

1–3, pinnately or palmately compound, 3(5)-foliolate, bases not auriculate;

terminal leaflets ovate, oblong, or linear; (0.5)1–3.5(6) cm;

lateral leaflets smaller; entire, dentate, or lobed.

(3)4–8, pinnately or palmately compound, 3(5)-foliolate, bases not auriculate, pubescent along margins;

terminal leaflets ovate to lanceolate, 2–7 × 0.6–4 cm.

Inflorescences

bracts 0, fruiting pedicels ascending to divaricate, 1–3.5 cm.

bracts 0, fruiting pedicels ascending to divaricate; (9)12–25 mm.

Flowers

sepals erect, oblong, 3.5–5 mm;

lateral pair saccate;

petals obovate, 10–15 × 4–7.5 mm, purple to pale pink, rarely white, not clawed;

tips rounded;

ovules 8–16 per ovary;

styles 4–8 mm.

sepals oblong, 2.5–4 mm;

lateral pair saccate;

petals white or rarely pinkish, obovate, 8–15 × 4–8 mm, clawed;

tips rounded or emarginate;

ovules 10–16 per ovary;

styles (0.5)1–4 mm.

Fruits

25–56 × 2–2.3 mm, glabrous.

15–32 × 1.4–2 mm, glabrous.

Seeds

oblong, 2–2.5 × 1.4–1.6 mm.

oblong, 1.8–2.3 × 1–1.2 mm.

2n

=40.

Cardamine nuttallii

Cardamine angulata

Distribution
[WildflowerSearch map]
[BONAP county map]
[WildflowerSearch map]
[BONAP county map]
Discussion

Damp woods, shaded bottomlands and hillsides, mossy slopes, streamsides. Flowering Mar–Jul. 0–2000 m. Casc, Col, CR, ECas, Est, Sisk, WV. CA, WA; north to British Columbia. Native.

Wet meadows, moist grounds, thickets, streambanks, swampy or damp woods. Flowering Apr–Jun. 0–1400 m. Casc, CR, Est, Sisk, WV. CA, WA; north to AK. Native.

Source Flora of Oregon, volume 2 draft
Ihsan Al-Shehbaz
Flora of Oregon, volume 2, page 464
Ihsan Al-Shehbaz
Sibling taxa
C. angulata, C. bellidifolia, C. breweri, C. californica, C. cordifolia, C. flexuosa, C. hirsuta, C. holmgrenii, C. occidentalis, C. oligosperma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica
C. bellidifolia, C. breweri, C. californica, C. cordifolia, C. flexuosa, C. hirsuta, C. holmgrenii, C. nuttallii, C. occidentalis, C. oligosperma, C. parviflora, C. pattersonii, C. penduliflora, C. pensylvanica
Synonyms Cardamine nuttallii var. covilleana, Cardamine nuttallii var. dissecta, Cardamine nuttallii var. gemmata, Cardamine nuttallii var. nuttallii, Cardamine pulcherrima, Cardamine pulcherrima var. pulcherrima, Cardamine pulcherrima var. tenella, Dentaria gemmata, Dentaria tenella, Dentaria tenella var. pulcherrima, Dentaria tenella var. quercetorum, Dentaria tenella var. tenella
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