Zizania palustris |
Poaceae subfam. ehrhartoideae |
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interior wildrice, northern wild rice, wild rice |
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Habit | Plants annual. | Plants annual or perennial. | ||||||||
Culms | to 3 m, erect, usually at least partly immersed. |
annual, sometimes woody, hollow or solid. |
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Sheaths | glabrous or with scattered hairs; ligules 3-16 mm, upper ligules truncate, lanceolate or triangular, erose; blades 20-60 cm long, 3-21(40+) mm wide, glabrous, margins glabrate or scabrous. |
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Leaves | distichous; sheaths open; auricles sometimes present; abaxial ligules absent; adaxial ligules membranous, scarious, or of hairs; pseudopetioles sometimes present; blades rarely cordate or sagittate at the base, venation parallel; mesophyll not radiate; adaxial palisade layer usually absent; fusoid cells sometimes present; arm cells absent or present; Kranz anatomy not developed; midribs simple or complex; adaxial bulliform cells present; stomates with dome-shaped or triangular subsidiary cells; bicellular microhairs present, terminal cells tapered; papillae sometimes present. |
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Panicles | 24-60 cm long, 1-20(40) cm wide; branches unisexual. |
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Inflorescences | panicles, racemes, or spikes, rarely with bracts other than those of the spikelets; disarticulation usually above the glumes, sometimes beneath the spikelets or at the base of the primary branches. |
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Spikelets | bisexual or unisexual, with 1 pistillate or bisexual floret, sometimes with 1-2 sterile florets below the functional floret. |
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Glumes | absent or 2; lemmas without uncinate hairs, sometimes terminally awned, awns single; paleas well-developed, lacking in sterile florets; lodicules 2, usually membranous, rarely fleshy, heavily vascularized; anthers (1)3-6(16); ovaries glabrous, without an apical appendage; styles 2, free to the base to fused throughout, 2-branched. |
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Caryopses | 6-30 mm long, 0.6-2 mm wide. |
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Fruits | caryopses or achenes; hila long-linear; endosperm without lipid, usually containing compound starch grains, rarely with simple starch grains; embryos to 1/3 the length of the caryopses; epiblasts usually present; scutellar cleft usually present; mesocotyl internode absent or very short; embryonic leaf margins usually overlapping. |
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Staminate | branches ascending or divergent; pedicel apices 0.2-0.4 mm wide.; staminate spikelets 6-17 mm, lanceolate, acuminate or awned, awns to 2 mm. |
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Pistillate | branches mostly appressed or ascending, a few sometimes divergent; pedicel apices 0.7-1.2 mm wide.; pistillate spikelets 8-33 mm long, 1-2.6 mm wide, lanceolate or oblong, coriaceous or indurate, lustrous, glabrous or with lines of short hairs, apices usually hirsute and abruptly narrowed, awned, awns to 10 cm; lemmas and paleas remaining clasped at maturity; aborted pistillate spikelets 0.6-2.6 mm wide. |
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x | =12 (10,15,17). |
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2n | = 30. |
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Zizania palustris |
Poaceae subfam. ehrhartoideae |
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Distribution |
AL; AR; AZ; CA; CO; CT; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; VT; WA; WI; WV; BC; MB; NB; NS; ON; PE; QC; SK
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Discussion | Zizania palustris grows mostly to the north of Z. aquatica, but the two species overlap in the Great Lakes region, eastern Canada, and New England. It is cultivated as a crop in some provinces and states, with California being the largest producer. All records from the western part of the Flora region reflect deliberate plantings; none are known to have persisted. In cultivated strains, the pistillate spikelets remain on the plant at maturity. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The Ehrhartoideae encompasses three tribes, one of which, the Oryzeae, is native to the Flora region; the Ehrharteae is represented by introduced species. The third tribe, Phyllorachideae C.E. Hubb., is native to Africa and Madagascar. It was included in the subfamily on the basis of its morphological similarity to the other two tribes. There are approximately 120 species in the Ehrhartoideae. They grow in forests, open hillsides, and aquatic habitats. Molecular data provide strong support for the close relationship of the Oryzeae and Ehrharteae (Grass Phylogeny Working Group 2001). Morphologically, they are characterized by spikelets that have a distal unisexual or bisexual floret with up to two proximal sterile florets, and the frequent presence of six stamens in the staminate or bisexual florets. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 24, p. 48. | FNA vol. 24, p. 32. | ||||||||
Parent taxa | Poaceae > subfam. Ehrhartoideae > tribe Oryzeae > Zizania | Poaceae | ||||||||
Sibling taxa | ||||||||||
Subordinate taxa | ||||||||||
Name authority | L. | Link | ||||||||
Web links |
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