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interior wildrice, northern wild rice, wild rice

Habit Plants annual. Plants annual or perennial.
Culms

to 3 m, erect, usually at least partly immersed.

annual, sometimes woody, hollow or solid.

Sheaths

glabrous or with scattered hairs;

ligules 3-16 mm, upper ligules truncate, lanceolate or triangular, erose;

blades 20-60 cm long, 3-21(40+) mm wide, glabrous, margins glabrate or scabrous.

Leaves

distichous;

sheaths open;

auricles sometimes present;

abaxial ligules absent;

adaxial ligules membranous, scarious, or of hairs;

pseudopetioles sometimes present;

blades rarely cordate or sagittate at the base, venation parallel;

mesophyll not radiate;

adaxial palisade layer usually absent;

fusoid cells sometimes present;

arm cells absent or present;

Kranz anatomy not developed;

midribs simple or complex;

adaxial bulliform cells present;

stomates with dome-shaped or triangular subsidiary cells;

bicellular microhairs present, terminal cells tapered;

papillae sometimes present.

Panicles

24-60 cm long, 1-20(40) cm wide;

branches unisexual.

Inflorescences

panicles, racemes, or spikes, rarely with bracts other than those of the spikelets;

disarticulation usually above the glumes, sometimes beneath the spikelets or at the base of the primary branches.

Spikelets

bisexual or unisexual, with 1 pistillate or bisexual floret, sometimes with 1-2 sterile florets below the functional floret.

Glumes

absent or 2;

lemmas without uncinate hairs, sometimes terminally awned, awns single;

paleas well-developed, lacking in sterile florets;

lodicules 2, usually membranous, rarely fleshy, heavily vascularized;

anthers (1)3-6(16);

ovaries glabrous, without an apical appendage;

styles 2, free to the base to fused throughout, 2-branched.

Caryopses

6-30 mm long, 0.6-2 mm wide.

Fruits

caryopses or achenes;

hila long-linear;

endosperm without lipid, usually containing compound starch grains, rarely with simple starch grains;

embryos to 1/3 the length of the caryopses;

epiblasts usually present;

scutellar cleft usually present;

mesocotyl internode absent or very short;

embryonic leaf margins usually overlapping.

Staminate

branches ascending or divergent;

pedicel apices 0.2-0.4 mm wide.; staminate spikelets 6-17 mm, lanceolate, acuminate or awned, awns to 2 mm.

Pistillate

branches mostly appressed or ascending, a few sometimes divergent;

pedicel apices 0.7-1.2 mm wide.; pistillate spikelets 8-33 mm long, 1-2.6 mm wide, lanceolate or oblong, coriaceous or indurate, lustrous, glabrous or with lines of short hairs, apices usually hirsute and abruptly narrowed, awned, awns to 10 cm;

lemmas and paleas remaining clasped at maturity; aborted pistillate spikelets 0.6-2.6 mm wide.

x

=12 (10,15,17).

2n

= 30.

Zizania palustris

Poaceae subfam. ehrhartoideae

Distribution
from FNA
AL; AR; AZ; CA; CO; CT; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MN; MO; MT; NC; ND; NE; NH; NJ; NY; OH; OR; PA; RI; SD; TN; VT; WA; WI; WV; BC; MB; NB; NS; ON; PE; QC; SK
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Discussion

Zizania palustris grows mostly to the north of Z. aquatica, but the two species overlap in the Great Lakes region, eastern Canada, and New England. It is cultivated as a crop in some provinces and states, with California being the largest producer. All records from the western part of the Flora region reflect deliberate plantings; none are known to have persisted. In cultivated strains, the pistillate spikelets remain on the plant at maturity.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

The Ehrhartoideae encompasses three tribes, one of which, the Oryzeae, is native to the Flora region; the Ehrharteae is represented by introduced species. The third tribe, Phyllorachideae C.E. Hubb., is native to Africa and Madagascar. It was included in the subfamily on the basis of its morphological similarity to the other two tribes. There are approximately 120 species in the Ehrhartoideae. They grow in forests, open hillsides, and aquatic habitats.

Molecular data provide strong support for the close relationship of the Oryzeae and Ehrharteae (Grass Phylogeny Working Group 2001). Morphologically, they are characterized by spikelets that have a distal unisexual or bisexual floret with up to two proximal sterile florets, and the frequent presence of six stamens in the staminate or bisexual florets.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Lower pistillate branches with 9-30 spikelets; pistillate part of the inflorescence 10-40 cm or more wide, the branches ascending to widely divergent; plants 1-3 m tall; blades 10-40+ mm wide
var. interior
1. Lower pistillate branches with 2-8 spikelets; pistillate part of the inflorescence 1-8(15) cm wide, the branches appressed or ascending, or a few branches somewhat divergent; plants to 2 m tall; blades 3-21 mm wide
var. palustris
1. Spikelets with 2 sterile florets below the functional floret, both well-developed, at least the upper sterile floret as long as or longer than the functional floret; glumes from 1/2 as long as the spikelets to exceeding the florets; culms not aerenchymatous; plants of dry to damp habitats
Ehrharteae
1. Spikelets with 0-2 sterile florets below the functional floret, when present, sterile florets 1/8 - 9/10 as long as the functional floret; glumes absent or highly reduced; culms aerenchymatous; plants of wet habitats
Oryzeae
Source FNA vol. 24, p. 48. FNA vol. 24, p. 32. Author: Grass Phylogeny Working Group;.
Parent taxa Poaceae > subfam. Ehrhartoideae > tribe Oryzeae > Zizania Poaceae
Sibling taxa
Z. aquatica, Z. latifolia, Z. texana
Subordinate taxa
Z. palustris var. interior, Z. palustris var. palustris
Name authority L. Link
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