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clotbur, lampourde épineuse, spiny clotbur, spiny cocklebur, spiny cockleburr

cocklebur, lampourde

Habit Plants 10–60(–120+) cm; nodal spines usually in pairs, simple or 2–3-partite, 15–30+ mm. Annuals (coarse), 10–200+ cm.
Stems

erect, branched.

Leaves

petioles 1–15(–25+) mm;

blades ± ovate to lanceolate or lance-linear, 4–8(–12+) × 1–3(–5+) cm, often pinnately 3(–7+)-lobed, abaxial faces gray to white, densely strigose.

cauline; mostly alternate (proximal 2–6 sometimes opposite);

petiolate;

blades lanceolate, linear, ovate, rounded-deltate, or suborbiculate, often ± palmately or pinnately lobed, ultimate margins entire or ± toothed, faces hirtellous or ± strigose, usually gland-dotted as well.

Staminate heads

involucres ± saucer-shaped, 3–5 mm diam.;

phyllaries 6–16+ in 1–2+ series, distinct to bases;

receptacles conic to columnar;

paleae spatulate to cuneiform or linear, membranous, distally ± villous or hirtellous;

florets 20–50+, corollas whitish, ± funnelform, lobes 5, erect or reflexed (filaments connate, anthers distinct or weakly coherent).

Pistillate heads

involucres ± ellipsoid, 2–5+ mm diam. at anthesis (6–20+ mm diam. at maturity);

phyllaries 30–75+ in 6–12+ series, outer 5–8 distinct, the rest (sometimes interpreted as paleae) proximally connate, their distinct tips mostly ± hooked (the distal 1–3 usually longer, stouter, and not hooked), the whole becoming a hard, prickly perigynium (a bur);

florets 2, corollas 0.

Heads

discoid, either pistillate (proximal) or functionally staminate (distal), in racemiform to spiciform arrays or borne singly (in axils).

Cypselae

(black) ± fusiform, enclosed in obovoid to ellipsoid, hard, prickly, 2-chambered burs;

pappi 0.

Burs

10–12(–15+) mm.

x

= 18.

2n

= 36.

Xanthium spinosum

Xanthium

Phenology Flowering Jul–Oct.
Habitat Damp or seasonally wet, alkaline soils, waste places, margins of agriculture
Elevation 10–1000 m (0–3300 ft)
Distribution
from FNA
AL; AZ; CA; CO; CT; DE; FL; GA; IA; ID; IL; IN; KS; KY; MA; MD; ME; MI; MO; MS; MT; NC; NE; NH; NJ; NM; NV; NY; OH; OR; PA; RI; SC; TN; TX; UT; VA; WA; WV; NB; ON; QC; SK; Mexico; Central America; South America; widely established in Old World
[WildflowerSearch map]
[BONAP county map]
from USDA
New World; introduced nearly worldwide
[BONAP county map]
Discussion

Some authors have contended that Xanthium spinosum originated in South America and is introduced and/or naturalized everywhere else that it is found.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Species 2–3 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Nodal spines 0; leaf blades suborbiculate to ± pentagonal or deltate
X. strumarium
1. Nodal spines (1–)3-lobed, 15–30+ mm; leaf blades ± lanceolate to ovate or lance-linear
X. spinosum
Source FNA vol. 21, p. 19. FNA vol. 21, p. 19. Author: John L. Strother.
Parent taxa Asteraceae > tribe Heliantheae > subtribe Ambrosiinae > Xanthium Asteraceae > tribe Heliantheae > subtribe Ambrosiinae
Sibling taxa
X. strumarium
Subordinate taxa
X. spinosum, X. strumarium
Synonyms X. ambrosioides, X. spinosum var. inerme
Name authority Linnaeus: Sp. Pl. 2: 987. (1753) Linnaeus: Sp. Pl. 2: 987. (1753): Gen. Pl. ed. 5, 424. (1754)
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