Woodsia scopulina subsp. appalachiana |
Woodsia scopulina subsp. laurentiana |
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laurent's cliff fern, Laurentian cliff fern, Laurentian Mountain cliff fern, Laurentian woodsia |
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Pinnae | with longest hairs composed of 5–8 cells. |
with longest hairs composed of 2–5 cells. |
Spores | averaging 39–46 µm. 2n = 76. |
averaging 50–57 µm. 2n = 152. |
Scales | of stems and petiole bases mostly bicolored with broad, usually continuous, dark central stripe, scales narrowly lanceolate. |
of stems and petiole bases (at least some) with clusters of dark, occluded cells near center forming narrow, usually discontinuous stripe, scales ovate-lanceolate. |
Indusial | segments broad, not at all filamentous. |
segments narrow, often filamentous distally. |
Woodsia scopulina subsp. appalachiana |
Woodsia scopulina subsp. laurentiana |
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Phenology | Sporulating summer–fall. | Sporulating summer–fall. |
Habitat | Shaded cracks and ledges on cliffs, mostly subacidic shale and sandstone | Cliffs and rocky slopes, found on a variety of substrates including both granite and limestone |
Elevation | 300–1700 m (1000–5600 ft) | 0–3000 m (0–9800 ft) |
Distribution |
AR; KY; NC; TN; VA; WV |
AZ; CA; CO; ID; MT; NV; OR; SD; UT; WA; WY; AB; BC; ON; QC |
Discussion | In addition to hybridizing with Woodsia scopulina subsp. scopulina (see comments above), subsp. laurentiana may have crossed with Woodsia oregana subsp. cathcartiana to form W. × maxonii R. M. Tryon. With very few collections and no biosystematic data available, however, the origin of this putative hybrid remains in doubt. Contrary to previous hypotheses (D. F. M. Brown 1964; D. B. Lellinger 1985), Great Lakes populations of W. scopulina were not involved in the origin of the local hybrid known as W. × abbeae (F. S. Wagner 1987). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | W. appalachiana, W. scopulina var. appalachiana | |
Name authority | (T. M. C. Taylor) Windham: Contr. Univ. Michigan Herb. 19: 58. (1993) | Windham: Contr. Univ. Michigan Herb. 19: 59. (1993) |
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