Woodsia obtusa subsp. obtusa |
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bluntlobe cliff fern |
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Stems | compact to short-creeping, individual branches usually 5–10 mm diam. |
Blade | coarsely cut and evidently 2-pinnate. |
Spores | averaging 42–47 µm. 2n = 152. |
Proximal | pinnules of lower pinnae usually shallowly lobed or merely dentate. |
Woodsia obtusa subsp. obtusa |
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Phenology | Sporulating summer–fall. |
Habitat | Cliffs and rocky slopes (rarely terrestrial), found on a variety of substrates including both granite and limestone |
Elevation | 0–1000 m (0–3300 ft) |
Distribution |
AL; AR; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; NC; NE; NH; NJ; NY; OH; OK; PA; RI; SC; TN; TX; VA; VT; WI; WV; ON; QC |
Discussion | D. F. M. Brown (1964) hypothesized that tetraploid Woodsia obtusa might be an autopolyploid derived from W. oregana. Recent isozyme and spore ornamentation studies indicate, however, that these species are not closely related, and the discovery of a diploid cytotype of W. obtusa suggests a different (albeit autopolyploid) origin for this taxon (M. D. Windham 1993). Tetraploid subsp. obtusa crosses with diploid subsp. occidentalis; the resulting triploids are sterile and have malformed spores. It also hybridizes with W. oregana subsp. cathcartiana to form the sterile tetraploid hybrid known as W. × kansana Brooks. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. |
Parent taxa | |
Sibling taxa | |
Name authority | (Swartz) Torrey |
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