Woodsia glabella |
Woodsia alpina |
|
---|---|---|
smooth cliff fern, smooth woodsia, woodsie glabre |
alpine cliff fern, alpine woodsia, northern cliff fern, woodsie alpine |
|
Stems | compact, erect to ascending, with cluster of persistent petiole bases of ± equal length; scales uniformly brown, lanceolate. |
compact, erect to ascending, with cluster of persistent petiole bases of ± equal length; scales uniformly brown, lanceolate. |
Leaves | 3.5–15 × 0.5–1.2 cm. |
2.5–20 × 0.5–2.5 cm. |
Petiole | green or straw-colored throughout, articulate above base at swollen node, somewhat pliable and resistant to shattering. |
reddish brown or dark purple when mature, articulate above base at swollen node, relatively brittle and easily shattered. |
Blade | linear to linear-lanceolate, pinnate-pinnatifid proximally, glabrous or with occasional sessile glands, never viscid; rachis glabrous. |
linear to narrowly lanceolate, usually pinnate-pinnatifid proximally, lacking glands, never viscid; rachis with widely scattered hairs and scales. |
Pinnae | ovate-lanceolate to deltate, longer than wide, abruptly tapered to a rounded or broadly acute apex; largest pinnae with 1–3 pairs of pinnules; abaxial surface with isolated hairs and linear scales, adaxial surface glabrous. |
|
Indusia | of narrow hairlike segments, these uniseriate throughout, composed of cells many times longer than wide, usually surpassing mature sporangia. |
of narrow, hairlike segments, these uniseriate throughout, composed of cells many times longer than wide, usually surpassing mature sporangia. |
Spores | averaging 39–45 µm. 2n = 78. |
averaging 46–53 µm. |
Proximal | pinnae fan-shaped, wider than long; distal pinnae ovate-lanceolate, longer than wide, abruptly tapered to a rounded or broadly acute apex; largest pinnae with 1–3 pairs of pinnules, abaxial and adaxial surfaces glabrous. |
|
Pinnules | entire or broadly crenate; margins nonlustrous, thin, lacking cilia or translucent projections. |
entire or broadly crenate; margins nonlustrous, thin, with occasional isolated cilia, lacking translucent projections. |
Vein | tips slightly (if at all) enlarged, barely visible adaxially. |
tips often enlarged to form whitish hydathodes visible adaxially. |
Woodsia glabella |
Woodsia alpina |
|
Phenology | Sporulating summer–early fall. | Sporulating summer–early fall. |
Habitat | Shaded cracks and ledges on cliffs, mostly calcareous rocks, especially limestone | Crevices and ledges on cliffs (occasionally on rocky slopes), mostly slaty and calcareous rocks |
Elevation | 0–1500 m (0–4900 ft) | 0–1500 m (0–4900 ft) |
Distribution |
AK; ME; MN; NH; NY; VT; AB; BC; MB; NB; NF; NS; NT; ON; QC; SK; YT; Greenland; n Eurasia
|
AK; ME; MI; MN; NH; NY; VT; BC; MB; NB; NF; NS; NT; ON; QC; SK; YT; Greenland; n Eurasia |
Discussion | Woodsia glabella is a well-marked species occasionally confused with narrow, glabrescent forms of W. alpina and W. oregana subsp. oregana. These taxa are readily distinguished from W. glabella by their petioles, which are reddish brown or dark purple near the base. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Isozyme studies confirm the longstanding hypothesis that Woodsia alpina is an allotetraploid derived from hybridization between W. glabella and W. ilvensis (see reticulogram). Considerable disagreement exists concerning the chromosome number of W. alpina, but 2n = 160 seems most likely, given the numbers reported for the two parental species. Hybrids between W. alpina and W. ilvensis have been reported from both Europe and North America. These morphologically intermediate triploids with malformed spores have been called W. × gracilis (Lawson) Butters. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 2. | FNA vol. 2. |
Parent taxa | ||
Sibling taxa | ||
Synonyms | W. alpina var. glabella, W. hyperborea var. glabella | Acrostichum alpinum, W. alpina var. bellii, W. bellii, W. hyperborea, W. ilvensis var. alpina |
Name authority | R. Brown ex Richardson: in Franklin, Narr. Journey Polar Sea 754. (1823) | (Bolton) Gray: Nat. Arr. Brit. Pl. 2: 17. (1822) |
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