Flora of North America species comparison

compact, erect to ascending, with a few persistent petiole bases of unequal lengths;

scales often uniformly brown but at least some bicolored with dark central stripe and pale brown margins, narrowly lanceolate.

5–25 × 1.5–6 cm.

light brown or straw-colored throughout when mature, occasionally darker at very base, not articulate above base, relatively brittle and easily shattered.

narrowly lanceolate to ovate, pinnate-pinnatifid to 2-pinnate proximally, sparsely to moderately glandular, never viscid;

glandular hairs with thin stalks and slightly expanded tips;

rachis with glandular hairs and occasional hairlike scales.

ovate-deltate to elliptic, longer than wide, abruptly tapered to a rounded or broadly acute apex, occasionally attenuate;

largest pinnae with 4–9 pairs of pinnules;

abaxial and adaxial surfaces glandular, lacking nonglandular hairs or scales.

of relatively broad segments;

segments multiseriate most of length, usually divided and uniseriate distally, composed of ± isodiametric cells, often surpassing mature sporangia.

averaging 43–49 µm. 2n = 152.

dentate, often shallowly lobed;

margins lustrous adaxially, usually thickened, lacking cilia but sparsely glandular, with occasional 1–2-celled translucent projections.

tips enlarged to form whitish hydathodes visible adaxially.