Waltheria indica |
Waltheria |
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basora-prieta, hierba del soldado, malva, malva blanca, malva del monte, sleepy morning, uhaloa |
waltheria |
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Habit | Plants branched or nearly simple, to 7–12 dm, indument variable, tomentose to short-woolly, sparsely to densely pubescent. | Subshrubs [herbs, shrubs], prostrate to erect, taprooted. | ||||||||
Stems | to 8 mm wide, nodes even or extended to 0.5 mm. |
unarmed, hairy, hairs stellate and/or simple, sometimes also glandular. |
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Leaves | stipules linear-triangular, 3–6 mm; petiole 2.5–5 cm × 1–4 mm; blade concolor or discolor, drying olive, ovate-oblong to lanceolate, to 7(–10) × 5 cm, base subcordate to obtuse, margins crenate-serrate, serrate, irregularly serrate, or crenate-dentate, apex narrowly rounded, obtuse, or subacute, surfaces tomentose or pubescent, trichome rays 0.4–1 mm, and sparsely hispidulous, slightly scabrous or not. |
petiolate; stipules deciduous, narrowly triangular or linear-triangular; blade: margins dentate, serrate, or crenate. |
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Inflorescences | subsessile, or on short pedunclelike lateral branches to 3.6 cm, glomerules 5–20 along stems, compact, often dense, sympodial, at times lax clusters with terminal flower and subtending dichasia; bracteoles free or with 1 adherent to stipule forming an elliptic 2-cuspidate foliole at glomerule base, bracts subequal, lanceolate or linear, 0.3–1.5 mm wide, apex acute, entire. |
axillary, cincinni in double or compound dichasial clusters, glomerulate, or paniculiform, 5–40-flowered; epicalyx absent; bracts 4, unilateral subtending floral pair, unequal or subequal. |
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Flowers | sessile or subsessile; calyx 4.2–setose apically; styles 0.9–1.7 × 0.1–0.3 mm, lateral or excentric, broader and stellate-hirsute distal to base; stigmas 12-branched, plumose (1.2–1.4 × 0.5–0.8 mm, column 0.9–1.2 mm, branches to 0.7 mm) or dissolute-penicillate (0.6–1.2 × 0.7–1.6 mm, column 0.4–0.7 mm, branches to 1.1 mm). |
sweet-smelling, sessile or subsessile, homostylous [mostly distylous]; sepals connate 1/2–2/3 length, nectary present; petals late-deciduous, pale yellow, bright yellow, or yellow-orange, usually darker in corolla throat, spatulate, clawed [rarely not clawed], lamina obovate, oblong, oblanceolate, or obtriangular; androgynophore absent or present, to 0.7 mm; stamens connate most of length [connate basally to entirely]; anthers 2-thecate, oblong or ovate; staminodes 0; ovary 1-locular, obovoid or obconic, stipitate or not; ovules 2 per locule; styles included, cylindric; stigmas 10–40-branched, apex slightly exserted above stamen apices and corolla. |
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Fruits | capsules, 1-locular, obconic, obovoid, or oblique, apically hairy, dehiscence partially or completely loculicidal. |
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Capsules | oblique, obconic, or obovoid, 2.1–3 × 1.5–2 mm, walls with endocarp corneous for 1+ mm from apex, sometimes merely membranous along one or both valve margins laterally; partially apically dehiscent to 2-valvate. |
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Seeds | dark brown, obovoid or obconic, 2–2.2 × 1.2–1.5 × 1–1.3 mm, smooth. |
1(or 2) per locule, brown or black, slightly laterally compressed, obovoid, obconic, or oblique, smooth or minutely granulate; endosperm present; cotyledons oblate [widely elliptic to elliptic, circular]. |
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x | = 5, 6, 7, 10, 13. |
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2n | = 24, 26 (India), 40 (Africa). |
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Waltheria indica |
Waltheria |
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Phenology | Flowering and fruiting year-round. | |||||||||
Habitat | Ruderal communities, disturbed scrublands, scrub hummocks, pine-oak sandhills, dunes, open savannas, dry coastal hummocks, sandy or shell soil | |||||||||
Elevation | 0–1000 m (0–3300 ft) | |||||||||
Distribution |
AL; AZ; FL; TX; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia
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sw United States; se United States; Mexico; Central America; South America; West Indies; Bermuda; Asia; Africa; Atlantic Islands; Indian Ocean Islands; Pacific Islands; Australia; tropical and subtropical areas |
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Discussion | Robert Brown (in J. H. Tuckey 1818) was the first to unequivocally synonymize the names Waltheria americana and W. indica published by Linnaeus in the same work. Brown adopted the name W. indica for the combined species and thus gave priority to that name. Waltheria indica in the narrow sense is globally in need of revision, being very complex and problematic. Heterostyly has been reported for some populations of W. indica in India (B. Bahadur et al. 1996). Flora area populations have the pistil subequal (0.2–0.5 mm longer) and more or less homostylous, or up to 1 mm longer than stamens, more like monomorphic pin floral morphology. Multiple introductions are apparent from the variability seen within the flora area. Apparent hybrids of W. indica with other species are found in Arizona, Florida, and Texas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Species ca. 60 (3 in the flora). Waltheria albicans Turczaninow and W. rotundifolia Schrank are found in Mexico, and apparent hybrids of them with W. indica in the broad sense occur in the United States. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 215. | FNA vol. 6, p. 212. | ||||||||
Parent taxa | Malvaceae > subfam. Byttnerioideae > Waltheria | Malvaceae > subfam. Byttnerioideae | ||||||||
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Subordinate taxa | ||||||||||
Synonyms | W. americana | |||||||||
Name authority | Linnaeus: Sp. Pl. 2: 673. (1753) | Linnaeus: Sp. Pl. 2: 673. (1753): Gen. Pl. ed. 5. 304. (1754) | ||||||||
Web links |