Viola trinervata
Viola praemorsa
3-nerve violet, desert pansy, Rainier or sagebrush or three-nerve violet, Rainier violet, sagebrush violet, three-nerve violet
Astoria violet, Canary violet, marsh violet, upland yellow violet, yellow montane violet
1–4, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous, from single, vertical, deep-seated caudex.
1–3(–5), prostrate, decumbent, or erect, leafy proximally and distally, glabrous or puberulent, on caudex from usually vertical, subligneous rhizome.
basal and cauline;
basal: 1–7, palmately compound, leaflets 3–5;
stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute;
petiole 4.5–15 cm, glabrous;
blade reniform or ovate to ± orbiculate, 2–5 × 2.5–5 cm, coriaceous, base tapered, leaflets cleft or dissected into 2–3 elliptic, lanceolate, or oblanceolate lobes 2–7 mm wide, margins usually entire, eciliate, apex acute, mucronulate, surfaces glabrous (± glaucous), abaxial surface usually with prominent vein parallel to each margin;
cauline similar to basal except: stipules lanceolate;
petiole 1–5.5 cm;
blade 1–3 × 2–4.5 cm.
basal and cauline;
basal: 1–5;
stipules adnate to proximal 1/3 of petiole, forming 2 narrow, linear-lanceolate wings, margins entire, apex of each wing free, acute;
petiole 2.6–19.2 cm, glabrous or densely puberulent;
blade usually elliptic to ovate, sometimes oblong-lanceolate to ± orbiculate, 1.7–14(–17) × 0.8–5.3(–6.7) cm, base attenuate to ± truncate or subcordate, often oblique, margins usually crenate, serrulate, or serrate, sometimes entire, ciliate or eciliate, apex acute to obtuse, surfaces glabrous or densely puberulent;
cauline similar to basal except: stipules lanceolate to ovate, margins entire or toothed, with or without gland-tipped projections, apex acute to acuminate;
petiole 1.3–16.2 cm, glabrous or puberulent;
blade 2.3–11(–14.8) × 1.1–3.6(–5.5) cm, length 1.1–6.5 times width.
1.1–7 cm, glabrous.
4.4–27 cm, glabrous or puberulent.
sepals lanceolate, margins eciliate, auricles 0–1 mm;
petals: upper 2 often overlapping, dark reddish violet on both surfaces, lower 3 lilac, rarely white, lateral 2 bearded, with yellow patch basally and reddish violet patch distal to yellow patch, lowest 9–15 mm with yellow patch, dark reddish violet-veined, spur yellow, gibbous, 0.6–1.5 mm;
style head bearded; cleistogamous flowers absent.
sepals lanceolate, margins eciliate, auricles 1–2 mm;
petals deep lemon-yellow adaxially, upper 2, and sometimes lateral 2, brownish purple abaxially, lower 3 brownish purple-veined, lateral 2 sparsely bearded, lowest 11–19 mm, spur yellow or pale green, gibbous, 0.5–3 mm;
style head bearded; cleistogamous flowers axillary.
ovoid, 7–12 mm, glabrous.
ellipsoid to oblong, 6–14 mm, glabrous or finely puberulent.
tan, 3.2–4.5 mm.
medium to dark brown or red-brown, 2–3 mm, elaiosome completely covering funiculus.
= 36, 48.
Viola trinervata
Viola praemorsa
In some populations of Viola trinervata the lower three petals are white with a yellow area proximally (V. B. Baird 1942).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Varieties 3 (3 in the flora).
Viola praemorsa is a complex group that appears to be evolving. Sympatric populations and the similarity of their flowers provide opportunities for hybridization and introgression (D. M. Fabijan et al. 1987). Some botanists have suggested that all taxa in the V. nuttallii complex, which includes V. bakeri, V. nuttallii, V. praemorsa, V. tomentosa, and V. vallicola, should be treated as varieties of V. nuttallii, as C. L. Hitchcock et al. (1955–1969, vol. 3) did, on the basis that these taxa are more or less sympatric in range, intergrade with one another, and possess no distinctive gross morphological features by which they can be consistently recognized. S. L. Welsh et al. (1987) treated V. praemorsa and V. linguaefolia Nuttall as synonymous with V. nuttallii, stating that V. nuttallii (and related taxa) are acaulescent to short-caulescent. Although some plants have short internodes, all plants in the V. nuttallii complex are caulescent. We acknowledge the complexity and taxonomic difficulties inherent in the V. nuttallii complex; patterns of morphology, cytology, and leaf flavonoid chemistry provide a basis for recognizing infrataxa. Fabijan et al. conducted the most extensive study of the complex to date; their treatment of V. praemorsa is followed here.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Basal and cauline leaf blade bases ± truncate to subcordate; basal leaf blades 1.7–5.9(–7.3) cm, length 1.1–3 times width; plants 5.5–21.5 cm. | var. praemorsa |
1. Basal and cauline leaf blade bases attenuate; basal leaf blades 2.7–17 cm, length 1.7–6.5 times width; plants 10.5–36.5 cm | → 2 |
2. Basal leaf blades (3.4–)5.3–17 cm, usually ovate to elliptic, sometimes oblong-lanceolate, length 1.8–6.5 times width; peduncles 5.9–27 cm. | var. flavovirens |
2. Basal leaf blades 2.7–8.5 cm, usually ovate to elliptic, sometimes oblong-lanceolate, length 1.7–3.4 times width; peduncles 5–15 cm. | var. linguifolia |
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