Viola trinervata |
Viola missouriensis |
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3-nerve violet, desert pansy, Rainier or sagebrush or three-nerve violet, Rainier violet, sagebrush violet, three-nerve violet |
Missouri violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–15 cm. | Plants perennial, acaulescent, not stoloniferous, 5–50 cm; rhizomes thick, fleshy. |
Stems | 1–4, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous, from single, vertical, deep-seated caudex. |
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Leaves | basal and cauline; basal: 1–7, palmately compound, leaflets 3–5; stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute; petiole 4.5–15 cm, glabrous; blade reniform or ovate to ± orbiculate, 2–5 × 2.5–5 cm, coriaceous, base tapered, leaflets cleft or dissected into 2–3 elliptic, lanceolate, or oblanceolate lobes 2–7 mm wide, margins usually entire, eciliate, apex acute, mucronulate, surfaces glabrous (± glaucous), abaxial surface usually with prominent vein parallel to each margin; cauline similar to basal except: stipules lanceolate; petiole 1–5.5 cm; blade 1–3 × 2–4.5 cm. |
basal, 1–8, ascending to erect; stipules linear-lanceolate to broadly lanceolate, margins entire, sometimes distally glandular, apex acute; petiole 5–20 cm, glabrous; blade green abaxially, unlobed, usually narrowly to broadly deltate, 1.5–12 × 1.5–10 cm, not fleshy, base cordate or broadly cordate to truncate, margins ± crenate to serrate, ciliate or eciliate, apex acute to acuminate, surfaces glabrous, rarely pubescent. |
Peduncles | 1.1–7 cm, glabrous. |
3–25 cm, usually glabrous. |
Flowers | sepals lanceolate, margins eciliate, auricles 0–1 mm; petals: upper 2 often overlapping, dark reddish violet on both surfaces, lower 3 lilac, rarely white, lateral 2 bearded, with yellow patch basally and reddish violet patch distal to yellow patch, lowest 9–15 mm with yellow patch, dark reddish violet-veined, spur yellow, gibbous, 0.6–1.5 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to dark blue-violet, lowest and sometimes lateral 2 purple-veined, lateral 2 bearded, lowest beardless, rarely lightly bearded, 15–25 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
Capsules | ovoid, 7–12 mm, glabrous. |
ellipsoid, 5–12 mm, glabrous. |
Seeds | tan, 3.2–4.5 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 54. |
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Viola trinervata |
Viola missouriensis |
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Phenology | Flowering Mar–May. | Flowering Mar–May. |
Habitat | Sagebrush flats, dry, rocky hillsides, usually in gravelly soil | Swamps, thickets, stream banks, alluvial woods |
Elevation | 400–1200 m (1300–3900 ft) | 50–2000 m (200–6600 ft) |
Distribution |
OR; WA
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AL; AR; DC; DE; FL; GA; IA; IL; IN; KS; LA; MD; MN; MO; MS; NC; NE; NJ; NM; OK; SC; SD; TN; TX; VA; WI
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Discussion | In some populations of Viola trinervata the lower three petals are white with a yellow area proximally (V. B. Baird 1942). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
What many have treated as Viola affinis, especially in the southern Gulf coastal states, is likely to be V. missouriensis. N. H. Russell (1965) considered the two as likely part of a species complex inhabiting alluvial woods and wet areas and exhibiting the typical deltate leaf blade shape. L. E. McKinney (1992) considered V. missouriensis a variety of V. sororia. Viola missouriensis appears to have a closer affinity to V. affinis, as Russell suggested; current evidence suggests maintaining V. missouriensis as a separate species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 161. | FNA vol. 6, p. 138. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. beckwithii var. trinervata | V. candidula, V. langloisii, V. lucidifolia, V. sororia var. missouriensis |
Name authority | (Howell) Howell ex A. Gray: Bot. Gaz. 11: 290. (1886) | Greene: Pittonia 4: 141. (1900) |
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