Viola trinervata |
Viola clauseniana |
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3-nerve violet, desert pansy, Rainier or sagebrush or three-nerve violet, Rainier violet, sagebrush violet, three-nerve violet |
Clausen's violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–15 cm. | Plants perennial, acaulescent, not stoloniferous, 3–20 cm; rhizome short, slender, fleshy. |
Stems | 1–4, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous, from single, vertical, deep-seated caudex. |
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Leaves | basal and cauline; basal: 1–7, palmately compound, leaflets 3–5; stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute; petiole 4.5–15 cm, glabrous; blade reniform or ovate to ± orbiculate, 2–5 × 2.5–5 cm, coriaceous, base tapered, leaflets cleft or dissected into 2–3 elliptic, lanceolate, or oblanceolate lobes 2–7 mm wide, margins usually entire, eciliate, apex acute, mucronulate, surfaces glabrous (± glaucous), abaxial surface usually with prominent vein parallel to each margin; cauline similar to basal except: stipules lanceolate; petiole 1–5.5 cm; blade 1–3 × 2–4.5 cm. |
basal, 2–10, often prostrate, sometimes ascending; stipules narrowly lanceolate, margins faintly glandular-toothed, apex acute or obtuse; petiole 5–11 cm, glabrous; blade unlobed, ± deltate, 3–5 × 4–5 cm, base ± truncate, margins serrate, ciliate or eciliate, apex obtuse, surfaces usually glabrous, sometimes sparsely pubescent abaxially. |
Peduncles | 1.1–7 cm, glabrous. |
8–14 cm, glabrous. |
Flowers | sepals lanceolate, margins eciliate, auricles 0–1 mm; petals: upper 2 often overlapping, dark reddish violet on both surfaces, lower 3 lilac, rarely white, lateral 2 bearded, with yellow patch basally and reddish violet patch distal to yellow patch, lowest 9–15 mm with yellow patch, dark reddish violet-veined, spur yellow, gibbous, 0.6–1.5 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate to ovate, margins eciliate or ciliate around auricles, auricles 1–2 mm; petals light violet on both surfaces, lower 3 purple-veined, all beardless, lowest petal 8–20 mm, spur white, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
Capsules | ovoid, 7–12 mm, glabrous. |
oblong, 8–10 mm, glabrous. |
Seeds | tan, 3.2–4.5 mm. |
dark brown to black, 1.2–2 mm. |
2n | = 44. |
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Viola trinervata |
Viola clauseniana |
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Phenology | Flowering Mar–May. | Flowering Apr–Jun. |
Habitat | Sagebrush flats, dry, rocky hillsides, usually in gravelly soil | Hanging gardens, seeps, springs, shady areas |
Elevation | 400–1200 m (1300–3900 ft) | 1600 m (5200 ft) |
Distribution |
OR; WA
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UT |
Discussion | In some populations of Viola trinervata the lower three petals are white with a yellow area proximally (V. B. Baird 1942). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Viola clauseniana is endemic to Zion National Park, Washington County. M. S. Baker reported that its seeds were minutely roughed, a characteristic not recorded for other Viola species. Viola clauseniana was originally thought to be closely related to the acaulescent blue violets, most notably V. nephrophylla (S. L. Welsh et al. 1987; L. E. McKinney 1992). After contemplating the 2n = 44 chromosome count obtained by J. Clausen (1964), H. E. Ballard (pers. comm.) suggested that V. clauseniana might be more closely related with the stemless white violets (for example, V. blanda) than with V. nephrophylla. Viola clauseniana was considered a distinct species by N. H. Holmgren (2005d) and T. Marcussen and T. Karlsson (2010). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 161. | FNA vol. 6, p. 128. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. beckwithii var. trinervata | |
Name authority | (Howell) Howell ex A. Gray: Bot. Gaz. 11: 290. (1886) | M. S. Baker: Madroño 4: 194. (1938) |
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