Viola trinervata |
Viola beckwithii |
|
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3-nerve violet, desert pansy, Rainier or sagebrush or three-nerve violet, Rainier violet, sagebrush violet, three-nerve violet |
Beckwith's or Great Basin violet, Beckwith's violet, Great Basin violet, sagebrush pansy, sagebrush violet |
|
Habit | Plants perennial, caulescent, not stoloniferous, 5–15 cm. | Plants perennial, caulescent, not stoloniferous, 2–22 cm. |
Stems | 1–4, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous, from single, vertical, deep-seated caudex. |
1–3, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous or usually puberulent, on single, short, vertical, deep-seated caudex. |
Leaves | basal and cauline; basal: 1–7, palmately compound, leaflets 3–5; stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute; petiole 4.5–15 cm, glabrous; blade reniform or ovate to ± orbiculate, 2–5 × 2.5–5 cm, coriaceous, base tapered, leaflets cleft or dissected into 2–3 elliptic, lanceolate, or oblanceolate lobes 2–7 mm wide, margins usually entire, eciliate, apex acute, mucronulate, surfaces glabrous (± glaucous), abaxial surface usually with prominent vein parallel to each margin; cauline similar to basal except: stipules lanceolate; petiole 1–5.5 cm; blade 1–3 × 2–4.5 cm. |
basal and cauline; basal: 1–6 per caudex, palmately compound, ± 2-ternate or 3-ternate, leaflets 3; stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute; petiole 2–10.5 cm, usually puberulent; blade ovate to deltate, 2.4–5 × 3.5–4.5 cm, base tapered, ultimate leaflets dissected into oblong, elliptic, lanceolate, or oblanceolate lobes 1–7 mm wide, lobe margins entire, usually ciliate, apex acute to obtuse, mucronulate, surfaces usually puberulent, seldom glabrous, abaxial surface without prominent vein parallel to each margin; cauline similar to basal except: stipules linear, apex acuminate; petiole 2–5.7 cm, usually puberulent, rarely glabrous; blade 1–2.7 × 1.5–3 cm. |
Peduncles | 1.1–7 cm, glabrous. |
1.5–10.6(–15.7) cm, usually puberulent, seldom glabrous. |
Flowers | sepals lanceolate, margins eciliate, auricles 0–1 mm; petals: upper 2 often overlapping, dark reddish violet on both surfaces, lower 3 lilac, rarely white, lateral 2 bearded, with yellow patch basally and reddish violet patch distal to yellow patch, lowest 9–15 mm with yellow patch, dark reddish violet-veined, spur yellow, gibbous, 0.6–1.5 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins eciliate, auricles 0.1–1 mm; petals dark reddish violet on both surfaces, lower 3 usually lilac, rarely white or whitish, lateral 2 bearded, with yellow patch basally, dark reddish violet-veined, lowest 10–22 mm, with yellow patch, dark reddish violet-veined, spur whitish or yellowish, tinged purple, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers absent. |
Capsules | ovoid, 7–12 mm, glabrous. |
oblong-ovoid, 7–12 mm, glabrous. |
Seeds | tan, 3.2–4.5 mm. |
brown, 3–4 mm. |
2n | = 24. |
|
Viola trinervata |
Viola beckwithii |
|
Phenology | Flowering Mar–May. | Flowering Mar–May. |
Habitat | Sagebrush flats, dry, rocky hillsides, usually in gravelly soil | Dry or moist places, among shrubs or beneath pines |
Elevation | 400–1200 m (1300–3900 ft) | 900–2700 m (3000–8900 ft) |
Distribution |
OR; WA
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CA; ID; NV; OR; UT
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Discussion | In some populations of Viola trinervata the lower three petals are white with a yellow area proximally (V. B. Baird 1942). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In some populations of Viola beckwithii, the three lower petals are white with a yellow area proximally (V. B. Baird 1942). Leaves have been described as palmately biternate or triternate (L. Abrams and R. S. Ferris 1923–1960, vol. 3), ternately decompound into linear segments (C. L. Hitchcock et al. 1955–1969, vol. 3), palmately three-parted then bipinnately parted into ultimate linear or spatulate segments (P. A. Munz 1959), and palmately about three times three-parted into linear or spatulate-linear segments (W. L. Jepson 1951). Some populations in northern California are nearly or completely glabrous, which M. S. Baker recognized as var. glabrata. Viola beckwithii is reported to hybridize with V. utahensis (G. Davidse 1976). Observed pollinators of V. beckwithii in Utah include Apis mellifera Linnaeus and Anthophora ursina Cresson (Davidse). When Cottam described Viola bonnevillensis, he suggested that it could be a hybrid between V. beckwithii and V. utahensis, and G. Davidse (1976) concurred. The type specimen of V. bonnevillensis (Cottam 7067, UT) was examined by R. J. Little. Because no similar forms are known to have been collected since 1939, it is presumed that this taxon is a hybrid. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 161. | FNA vol. 6, p. 122. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. beckwithii var. trinervata | V. beckwithii var. cachensis, V. beckwithii subsp. glabrata, V. bonnevillensis |
Name authority | (Howell) Howell ex A. Gray: Bot. Gaz. 11: 290. (1886) | Torrey: in E. M. Durand, Pl. Pratten. Calif., 82. (1855) |
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