Viola trinervata |
Viola adunca |
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3-nerve violet, desert pansy, Rainier or sagebrush or three-nerve violet, Rainier violet, sagebrush violet, three-nerve violet |
Cascades early blue violet, common periwinkle, early blue violet, hook-spur violet, hookedspur violet, large periwinkle, sand violet, western dog violet, wild dog violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–15 cm. | Plants perennial, caulescent, not stoloniferous, 1.8–30(–35) cm. | ||||
Stems | 1–4, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous, from single, vertical, deep-seated caudex. |
1–5, erect, ascending, or decumbent, sometimes later reclining to nearly prostrate, glabrous or puberulent, on caudex from subligneous rhizome. |
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Leaves | basal and cauline; basal: 1–7, palmately compound, leaflets 3–5; stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute; petiole 4.5–15 cm, glabrous; blade reniform or ovate to ± orbiculate, 2–5 × 2.5–5 cm, coriaceous, base tapered, leaflets cleft or dissected into 2–3 elliptic, lanceolate, or oblanceolate lobes 2–7 mm wide, margins usually entire, eciliate, apex acute, mucronulate, surfaces glabrous (± glaucous), abaxial surface usually with prominent vein parallel to each margin; cauline similar to basal except: stipules lanceolate; petiole 1–5.5 cm; blade 1–3 × 2–4.5 cm. |
basal and cauline; basal: 1–4; stipules linear to linear-lanceolate, margins entire or laciniate with gland-tipped projections, apex acute to acuminate; petiole 0.5–13.5 cm, glabrous or puberulent; blade usually ovate or ovate-deltate to ovate-orbiculate, sometimes ± reniform or oblong, 0.5–6.9 × 0.4–5 cm, base cordate, subcordate, truncate, or attenuate, usually decurrent on petiole, margins crenate to crenulate or entire, ciliate or eciliate, apex acute to obtuse, surfaces glabrous or sparsely to densely puberulent; cauline similar to basal except: stipule margins lacerate to laciniate; petiole 0.5–6.5 cm; blade 0.6–5.5 × 0.4–4.7 cm. |
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Peduncles | 1.1–7 cm, glabrous. |
1.7–13.8 cm, glabrous or puberulent. |
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Flowers | sepals lanceolate, margins eciliate, auricles 0–1 mm; petals: upper 2 often overlapping, dark reddish violet on both surfaces, lower 3 lilac, rarely white, lateral 2 bearded, with yellow patch basally and reddish violet patch distal to yellow patch, lowest 9–15 mm with yellow patch, dark reddish violet-veined, spur yellow, gibbous, 0.6–1.5 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins ciliate or eciliate, auricles not enlarged in fruit, 0.5–2 mm; petals light- to deep- to lavender-violet on both surfaces, rarely white, lower 3 usually white basally, dark violet-veined, lateral 2 (and sometimes upper 2) bearded, lowest 7–17(–23) mm, spur purple to violet or white, elongated, 5–7 mm, tip straight or pointed, curved up or lateral; style head sparsely to densely bearded, sometimes beardless; cleistogamous flowers axillary. |
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Capsules | ovoid, 7–12 mm, glabrous. |
short-ovoid, 6–11 mm, glabrous. |
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Seeds | tan, 3.2–4.5 mm. |
dark brown to olive-black, 1.5–2 mm. |
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Viola trinervata |
Viola adunca |
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Phenology | Flowering Mar–May. | |||||
Habitat | Sagebrush flats, dry, rocky hillsides, usually in gravelly soil | |||||
Elevation | 400–1200 m (1300–3900 ft) | |||||
Distribution |
OR; WA
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AK; AZ; CA; CO; IA; ID; MA; ME; MI; MN; MT; ND; NE; NH; NM; NV; NY; OR; SD; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; NT; NU; ON; QC; SK; YT
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Discussion | In some populations of Viola trinervata the lower three petals are white with a yellow area proximally (V. B. Baird 1942). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Viola adunca is polymorphic with over 50 named taxa. G. D. McPherson and J. G. Packer (1974) reported that diploid, triploid, and tetraploid races of V. adunca occur in Canada and northwestern United States. They found that diploid and tetraploid chromosome races can be distinguished morphologically based on style beards and on the size of guard cells and pollen grains and recommended taxonomic recognition of both races. In diploid races, the style projections are more or less cylindrical and about one-sixth the width of the style; in tetraploid races they are short-conical or globular, and about one-tenth or less than the width of the style beard. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 161. | FNA vol. 6, p. 119. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. beckwithii var. trinervata | Lophion aduncum, V. canina var. adunca | ||||
Name authority | (Howell) Howell ex A. Gray: Bot. Gaz. 11: 290. (1886) | Smith: in A. Rees, Cycl. 37: Viola no. 63 | ||||
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