Viola tomentosa
Viola missouriensis
felt-leaf violet, woolly or felt-leaf or feltleaf violet, woolly violet
Missouri violet
1–3(–5), prostrate or decumbent to erect, leafy proximally and distally, densely white-tomentose, from usually vertical, subligneous rhizome.
basal and cauline;
basal: 1–6;
stipules linear to broadly ovate-oblong, margins entire, sometimes with scattered glandular hairs, apex acute to obtuse;
petiole 2–6 cm, densely white-tomentose;
blade ± oblanceolate to elliptic, 1.5–5 × 1.4–2.1 cm, base attenuate, usually oblique, margins usually entire, rarely crenate distally, ciliate, apex acute to usually obtuse, mucronulate, surfaces densely white-tomentose abaxially, strigose adaxially;
cauline similar to basal except: stipules ovate, lanceolate, oblanceolate, or oblong, margins entire or toothed, densely ciliate with white hairs;
petiole 1.5–3.5 cm;
blade 1.8–4 × 0.6–1.1 cm.
basal, 1–8, ascending to erect;
stipules linear-lanceolate to broadly lanceolate, margins entire, sometimes distally glandular, apex acute;
petiole 5–20 cm, glabrous;
blade green abaxially, unlobed, usually narrowly to broadly deltate, 1.5–12 × 1.5–10 cm, not fleshy, base cordate or broadly cordate to truncate, margins ± crenate to serrate, ciliate or eciliate, apex acute to acuminate, surfaces glabrous, rarely pubescent.
1–4 cm, densely white-tomentose.
3–25 cm, usually glabrous.
sepals lanceolate, margins ciliate-tomentose, auricles 0.5–1 mm;
petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 bearded, lowest 6–11 mm, spur yellow, gibbous, 0.5–1.5 mm;
style head bearded; cleistogamous flowers absent.
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm;
petals light to dark blue-violet, lowest and sometimes lateral 2 purple-veined, lateral 2 bearded, lowest beardless, rarely lightly bearded, 15–25 mm, spur same color as petals, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on prostrate to ascending peduncles.
± spherical, 4–5 mm, densely white-tomentose.
ellipsoid, 5–12 mm, glabrous.
brown with lighter brown mottling, 2.5–2.8 mm.
beige, mottled to bronze, 1.5–2.5 mm.
= 12.
= 54.
Viola tomentosa
Viola missouriensis
Viola tomentosa occurs in El Dorado, Nevada, Placer, Plumas, and Sierra counties. M. S. Baker (1949) reported that nearly every leaf axil of V. tomentosa produces a flower bud and that these buds produce chasmogamous flowers late in season instead of cleistogamous flowers, as do other members of the V. nuttallii complex.
Viola tomentosa hybridizes with V. purpurea; the hybrids appeared sterile (M. S. Baker 1949). J. Clausen (1964) reported a putative hybrid between V. tomentosa and V. sheltonii from one location in Sierra County.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
What many have treated as Viola affinis, especially in the southern Gulf coastal states, is likely to be V. missouriensis. N. H. Russell (1965) considered the two as likely part of a species complex inhabiting alluvial woods and wet areas and exhibiting the typical deltate leaf blade shape. L. E. McKinney (1992) considered V. missouriensis a variety of V. sororia. Viola missouriensis appears to have a closer affinity to V. affinis, as Russell suggested; current evidence suggests maintaining V. missouriensis as a separate species.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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