Viola tomentosa |
Viola lithion |
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felt-leaf violet, woolly or felt-leaf or feltleaf violet, woolly violet |
rock violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 7–10 cm. | Plants perennial, caulescent, not stoloniferous, 5–15 cm. |
Stems | 1–3(–5), prostrate or decumbent to erect, leafy proximally and distally, densely white-tomentose, from usually vertical, subligneous rhizome. |
1–3, ascending to erect, glabrous, on caudex from fleshy rhizome. |
Leaves | basal and cauline; basal: 1–6; stipules linear to broadly ovate-oblong, margins entire, sometimes with scattered glandular hairs, apex acute to obtuse; petiole 2–6 cm, densely white-tomentose; blade ± oblanceolate to elliptic, 1.5–5 × 1.4–2.1 cm, base attenuate, usually oblique, margins usually entire, rarely crenate distally, ciliate, apex acute to usually obtuse, mucronulate, surfaces densely white-tomentose abaxially, strigose adaxially; cauline similar to basal except: stipules ovate, lanceolate, oblanceolate, or oblong, margins entire or toothed, densely ciliate with white hairs; petiole 1.5–3.5 cm; blade 1.8–4 × 0.6–1.1 cm. |
basal and cauline; basal: 1–4; stipules unknown; petiole 1–10 cm, glabrous, sometimes finely puberulent; blade broadly ovate or deltate to broadly deltate, 1–2.5(–2.9) × 0.6–2.2(–2.6) cm, base usually cordate to truncate, rarely rounded, margins crenate-dentate, eciliate, apex acute, surfaces glabrous; cauline similar to basal except: stipules lanceolate, margins usually fimbriate-toothed, sometimes entire, apex attenuate or acute; petiole 1.1–3.7 cm, sometimes finely puberulent; blade ovate to deltate, 0.7–2.2 × 0.4–1.2 cm, base sometimes rounded on distal blades. |
Peduncles | 1–4 cm, densely white-tomentose. |
3–6(–10) cm, glabrous. |
Flowers | sepals lanceolate, margins ciliate-tomentose, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 often brownish purple abaxially, lower 3 dark brown- to brownish purple-veined, lateral 2 bearded, lowest 6–11 mm, spur yellow, gibbous, 0.5–1.5 mm; style head bearded; cleistogamous flowers absent. |
sepals lanceolate, margins eciliate, auricles 0.5–1 mm; petals blue to pale violet on both surfaces with yellow area basally, lower 3 purple-veined, lowest with yellow area, lateral 2 bearded, lowest 5.5–11 mm, spur white to pale violet, gibbous, 0.5–1.3 mm; style head bearded; cleistogamous flowers unknown. |
Capsules | ± spherical, 4–5 mm, densely white-tomentose. |
subglobose, ca. 5 mm, glabrous. |
Seeds | brown with lighter brown mottling, 2.5–2.8 mm. |
dark brown, ca. 1.8 mm. |
2n | = 12. |
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Viola tomentosa |
Viola lithion |
|
Phenology | Flowering May–Aug. | Flowering Jun–Aug. |
Habitat | Dry, gravelly places, open ponderosa, Jeffrey, lodgepole pine forests | Seasonally wet cracks and crevices, narrow ledges of rock outcrops, shaded northeast-facing avalanche chutes, cirque headwalls, subalpine conifer zone |
Elevation | 1500–2000 m (4900–6600 ft) | 2300–3100 m (7500–10200 ft) |
Distribution |
CA
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NV; UT |
Discussion | Viola tomentosa occurs in El Dorado, Nevada, Placer, Plumas, and Sierra counties. M. S. Baker (1949) reported that nearly every leaf axil of V. tomentosa produces a flower bud and that these buds produce chasmogamous flowers late in season instead of cleistogamous flowers, as do other members of the V. nuttallii complex. Viola tomentosa hybridizes with V. purpurea; the hybrids appeared sterile (M. S. Baker 1949). J. Clausen (1964) reported a putative hybrid between V. tomentosa and V. sheltonii from one location in Sierra County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Viola lithion is known only from the White Pine Range in Nevada and the Pilot Range straddling the Nevada-Utah border. It is related to V. canadensis and V. flettii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 160. | FNA vol. 6, p. 136. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Name authority | M. S. Baker & J. C. Clausen: Leafl. W. Bot. 5: 142. (1949) | N. H. Holmgren & P. K. Holmgren: Brittonia 44: 300, fig. 1A – D. (1992) |
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