Viola striata |
Viola brittoniana |
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cream or cream-white or pale or striped or striped cream violet, creamy violet, striped cream violet, striped white violet, violette strièe |
coast violet, coastal violet, northern coastal violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 10–60 cm. | Plants perennial, acaulescent, not stoloniferous, 5–30 cm; rhizome thick, fleshy. |
Stems | 1–4, ascending to erect (often declining during anthesis), glabrous or pubescent, on caudex from fleshy rhizome. |
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Leaves | basal and cauline; basal: 2–6; stipules lanceolate to narrowly deltate, margins laciniate, apex acute; petiole 3–6 cm, glabrous or puberulent; blade ovate to reniform, 2–7 × 1–2.5 cm, base cordate, margins crenate to serrate, ciliate or eciliate, apex obtuse to acute, surfaces glabrous or pubescent; cauline similar to basal except: stipules lanceolate, margins laciniate; petiole 3–7 cm; distal blades ovate to deltate, 1–6 × 1–4 cm, base cordate, apex acuminate to acute. |
basal, 5–9, ascending to erect, 5–9-lobed; stipules linear-lanceolate, margins entire, apex obtuse; petiole 3–16 cm, usually glabrous; mid-season blades incised or lobed throughout, earliest blades lobed (plants homophyllous), similar to mid-season blades, middle and lateral blade lobes differ in width and/or shape, middle lobes lanceolate or spatulate to narrowly obovate, lateral lobes lanceolate or spatulate to falcate (each sometimes with deltate to falcate appendages or teeth along margins), 1–7 × 2–8 cm, base truncate to cordate, margins entire, ciliate, apex acute to obtuse, surfaces usually glabrous, rarely with a few strigose, hairs concentrated on veins. |
Peduncles | 5–12 cm, glabrous or pubescent. |
5–18 cm, usually glabrous. |
Flowers | sepals lanceolate, margins ciliate or eciliate, auricles 2–3 mm; petals white or cream on both surfaces, without yellow patch basally, lowest and usually lateral 2 purple-veined, lateral 2 and sometimes all densely bearded, lowest 10–18 mm, spur white, gibbous to elongated, 3–6 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 2–3 mm; petals light to soft reddish violet on both surfaces, lower 3 white basally, dark violet-veined, lateral 2 bearded, lowest 10–25 mm, sometimes bearded, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on ascending to erect peduncles. |
Capsules | ellipsoid, 6–7 mm, glabrous. |
ellipsoid, 10–15 mm, glabrous. |
Seeds | beige to bronze, 1.5–3 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 20. |
= 54. |
Viola striata |
Viola brittoniana |
|
Phenology | Flowering Mar–Jun. | Flowering Apr–Jun. |
Habitat | Riparian or alluvial woods, floodplains in silty loam, meadows | Alluvial woods, mesic forests on slopes near streams, wet fields, salt meadows |
Elevation | 40–1000 m (100–3300 ft) | 0–100 m (0–300 ft) |
Distribution |
AL; AR; CT; DC; DE; GA; IA; IL; IN; KY; MA; MD; MI; MO; NC; NH; NJ; NY; OH; OK; PA; SC; TN; VA; WI; WV; ON
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CT; DE; MA; MD; ME; NC; NJ; NY; PA; SC; VA |
Discussion | Dead, long-persistent stems of Viola striata are often present the following season. In flower, the plants are often mistaken for V. canadensis because the petals are whitish (H. E. Ballard 1992). Viola striata reportedly hybridizes with V. labradorica (= V. ×eclipes H. E. Ballard), V. rostrata (= V. ×brauniae Grover ex Cooperrider), V. walteri var. appalachiensis (= V. ×wujekii H. E. Ballard), and V. walteri var. walteri (= V. ×cooperrideri H. E. Ballard). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Britton considered Viola brittoniana similar to V. pedatifida, apparently making him the first to recognize their affinity. Both are homophyllous and similar in other characters (L. E. McKinney 1992). McKinney recognized V. brittoniana as a variety of V. pedatifida. Additional evidence (N. L. Gil-Ad 1997; A. Haines 2011) suggests that these taxa are sufficiently distinct to continue recognition as separate species. Viola pectinata has sharply dentate leaves and is closely related to V. brittoniana, usually occurring with it. N. L. Gil-Ad (1997) made a convincing argument to recognize it as a form; A. Haines (2011) recognized it as a species. Others have considered it to be a sporadic form that may be of hybrid origin, or, as N. H. Russell (1965) suggested, is due to genetic dimorphism. We are reserving a decision on treating this taxon until additional studies are completed. A. Haines (2011) reported that the distribution of Viola brittoniana in Maine is actually based on the hybrid V. ×insolita House (V. brittoniana × V. sororia). Viola brittoniana reportedly hybridizes with V. cucullata (= V. ×notabilis E. P. Bicknell), V. sagittata var. sagittata (= V. ×mulfordiae Pollard), and V. sororia (= V. ×insolita). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 159. | FNA vol. 6, p. 125. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | Lophion striatum, V. conspersa var. masonii | V. atlantica, V. baxteri, V. brittoniana var. pectinata, V. pectinata, V. pedatifida subsp. brittoniana, V. pedatifida var. brittoniana |
Name authority | Aiton: Hort. Kew. 3: 290. (1789) | Pollard: Bot. Gaz. 26: 332. (1898) |
Web links |