Viola sororia
Viola subsinuata
common blue violet, common or downy or woolly blue violet, northern blue violet, northern violet, northern woodland violet, violette parente, woolly blue violet
early blue violet, wavy-leaf violet, wood violet
basal, 1–8, ascending to erect;
stipules linear-lanceolate to broadly lanceolate, margins entire, sometimes glandular distally, apex acute;
petiole 2–25 cm, pubescent or glabrous;
blade green abaxially, unlobed, ovate or broadly ovate to reniform, 2–5 × 2–10 cm, not fleshy, base cordate, margins crenate to serrate, ciliate or eciliate, apex acute to obtuse or rounded, surfaces usually pubescent, rarely glabrous.
basal, 2–11, ascending to erect;
stipules linear-lanceolate, margins entire, apex acute;
petiole 5–25 cm, glabrous or pubescent;
mid-season leaf blades incised or lobed throughout, earliest leaf blades lobed (plants homophyllous), similar to mid-season blades, blade 5–9(–16)-lobed, sinuses usually narrower, shallower toward leaf base, middle and lateral blade lobes differ in width and/or shape, middle lobes narrowly deltate to narrowly elliptic, lateral lobes narrowly elliptic, lanceolate, or falcate, 1–11 × 1–12 cm, base truncate to cordate, margins entire or crenate, ciliate or eciliate, apex acute, mucronulate, surfaces usually pubescent throughout or on veins.
3–25 cm, glabrous or sparsely pubescent.
5–15 cm, glabrous or pubescent.
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm;
petals light to dark blue- or dark purple-violet, reddish purple, or rarely white on both surfaces, usually white basally, lowest and sometimes lateral 2 purple-veined, lateral 2 bearded, lowest bearded or beardless, 15–25 mm, spur same color as petals, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on prostrate to ascending peduncles.
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm;
petals light to dark blue-violet on both surfaces, lower 3 and upper 2 sometimes purple-veined, lateral 2 bearded, lowest sometimes bearded, 15–25 mm, spur color same as petals, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on prostrate to ascending peduncles.
ellipsoid, 5–12 mm, glabrous.
ellipsoid, 8–12 mm, glabrous.
beige, mottled to bronze, 1.5–2.5 mm.
beige, mottled to bronze, 1.5–2.5 mm.
= 54.
= 54.
Viola sororia
Viola subsinuata
Viola sororia is similar to V. palmata in the high degree of phenotypic plasticity. The petal color is usually light to dark blue-violet. In V. sororia forma priceana (the Confederate violet) petals are grayish white with violet veins. Viola septentrionalis was recognized by N. L. Gil-Ad (1997) based on what he considered distinct capsule and seed morphology. His assumptions were based on one to four specimens. A. Haines (2011) alluded to the variability of V. sororia and indicated that a more northern form has often been called V. septentrionalis. He suggested that range-wide patterns of variation make it difficult or impractical to separate. We currently see no valid reason to recognize V. septentrionalis.
Viola floridana was recognized by D. B. Ward (2006) as distinct from V. sororia based on being glabrous except for scattered hairs on petioles.
Viola sororia reportedly hybridizes with V. cucullata (= V. ×bissellii House), V. hirsutula [= V. ×cordifolia (Nuttall) Schweinitz], V. pedatifida var. brittoniana (= V. ×insolita House), V. pedatifida var. pedatifida (= V. ×bernardii Greene), and V. sagittata var. sagittata (= V. ×conjugens Greene).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
L. E. McKinney (1992) described the misconceptions surrounding Viola subsinuata that many came to think of as the heterophyllous V. palmata, whereas V. subsinuata is homophyllous. As E. Brainerd (1910, 1921) pointed out, heterophylly versus homophylly is an important and steadfast character difference in acaulescent Viola species. N. L. Gil-Ad (1997) chose not to recognize V. subsinuata, suggesting that it represented a hybrid or introgressant. H. E. Ballard (2000) and A. Haines (2011) both recognized V. subsinuata as a species.
Viola subsinuata is a state historical species in Rhode Island, where it was last documented in 1941 (R. W. Enser, http://rinhs.org/wp-content/uploads/ri_rare_plants_2007.pdf).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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