Viola sempervirens |
Viola palustris |
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evergreen violet, redwood violet, redwoods violet, trailing yellow violet, violette toujours verte |
alpine marsh violet, marsh violet, northern marsh or marsh or swamp or alpine marsh violet, northern marsh violet, violette des marais |
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Habit | Plants perennial, caulescent, stoloniferous, 10–30 cm; stolons green or reddish, leafy, sometimes rooting at nodes, becoming lignified in age. | Plants perennial, acaulescent, stoloniferous, 3–21.5 cm; stolons pale, often rooting and leafy at nodes; rhizome slender, fleshy. |
Stems | 1–5, prostrate, spreading, glabrous or sparsely puberulent, from current and/or previous year’s growth, on usually vertical, fleshy rhizome, rooting and forming rosettes at or near tip; rooted rosettes often develop into an erect, fleshy caudex from which new stems are produced. |
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Leaves | evergreen, basal and cauline; basal: 1–6(–10); stipules deltate to ovate or linear-lanceolate, margins entire or glandular-toothed, apex acute to long-acuminate; petiole 2–16 cm, glabrous; blade often purple-spotted abaxially and/or adaxially, orbiculate to ovate, 1–4.5 × 2–3.9 cm, base cordate to truncate, margins crenate, eciliate, apex blunt to obtuse, mucronulate, surfaces glabrous or with scattered bristles on one or both surfaces; cauline similar to basal except: stipules deltate to lanceolate, margins entire or sparingly toothed; petiole 0.3–3 cm; blade 1.2–2.2 × 1.2–2 cm. |
basal, 2–4, ascending to erect; stipules linear-lanceolate, margins entire, apex acute; petiole 1–17 cm, glabrous; blade unlobed, reniform or ovate to orbiculate, 0.5–6.4 × 0.5–5.5 cm, ± as long as wide, base cordate, margins crenulate, eciliate, apex obtuse to acuminate, surfaces glabrous. |
Peduncles | 5–10 cm, glabrous. |
2–20.7 cm, glabrous, bracteoles usually below middle in chasmogamous flowers. |
Flowers | sepals lanceolate to ovate-lanceolate, margins eciliate, auricles 1–2 mm; petals lemon-yellow on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded, lowest 8–17 mm, spur yellow or whitish, gibbous, 1–2.5 mm; style head bearded; cleistogamous flowers axillary. |
sepals ovate or lanceolate, margins eciliate, auricles 1–2 mm; petals lilac, pale blue, or pale violet on both surfaces, sometimes nearly white and tinged with lilac, lowest and sometimes lateral 2 dark violet-veined, lateral 2 bearded or beardless, lowest 8–16 mm, spur usually same color as petals, gibbous, 1–3 mm; style head beardless; cleistogamous flowers on ascending peduncles. |
Capsules | mottled with purple, spherical to ovoid, 5–8 mm, glabrous. |
ellipsoid, 6–10 mm, glabrous. |
Seeds | brown, tinged purple, 2–2.5 mm. |
dark brown, 1–2 mm. |
2n | = 24, 48. |
= 48. |
Viola sempervirens |
Viola palustris |
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Phenology | Flowering Jan–Jul. | Flowering Apr–Jul. |
Habitat | Redwood forests, other coastal forests, Douglas fir, other coniferous forests | Marshes, swamps, fens, wet forests, stream banks, brushy places, lowlands to subalpine zones |
Elevation | 30–1400 m (100–4600 ft) | 0–1800 m (0–5900 ft) |
Distribution |
AK; CA; ID; OR; WA; BC
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CA; CO; ID; ME; MT; NH; NV; OR; SD; UT; WA; WY; AB; BC; MB; NL; NT; NU; ON; QC; SK; Greenland; Eurasia
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Discussion | In California, Viola sempervirens occurs in shaded redwood forests and other coastal forest habitats. In Oregon and Washington, it occurs in Douglas fir and other coniferous forests, where it can form mats (clones) one meter or more in diameter; its prostrate, spreading growth habit is similar to V. walteri. The leafy stems of V. sempervirens are similar to the leafy stolons of V. odorata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola palustris occurs in small colonies; individual plants are interconnected by stolons. When flowering, usually three or more leaves are present per caudex. M. Sorsa (1968) noted that in western North America Viola palustris specimens differ from European V. palustris plants and often show variation in character combinations. She stated (pers. comm.) that North American V. palustris is not a uniform entity. E. Hultén (1968) synonymized Viola palustris with V. epipsila, as did W. J. Cody (2000). G. W. Douglas et al. (1998–2002, vol. 5) recognized both species. We assume that V. palustris does not occur in Alaska; verification is needed. See additional discussion under 18a. V. epipsila var. repens. M. Sorsa (1968) indicated that Viola palustris var. brevipes has often been confused with V. blanda and that var. brevipes may have arisen from hybridization between a tetraploid and a diploid plant. Listed here in synonymy, var. brevipes may warrant recognition. Reports of Viola palustris from one location in Coconino County, Arizona, first collected in 1962, likely refer to V. macloskeyi. Viola palustris hybridizes with V. epipsila var. repens (M. Sorsa 1968; T. Marcussen and T. Karlsson 2010). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 157. | FNA vol. 6, p. 142. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. sarmentosa, V. sempervirens subsp. orbiculoides | V. palustris subsp. brevipes, V. palustris var. brevipes, V. palustris var. leimonia, V. palustris var. pensylvanica, V. palustris subsp. pubifolia |
Name authority | Greene: Pittonia 4: 8. (1899) | Linnaeus: Sp. Pl. 2: 934. (1753) |
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