Viola sempervirens |
Viola beckwithii |
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evergreen violet, redwood violet, redwoods violet, trailing yellow violet, violette toujours verte |
Beckwith's or Great Basin violet, Beckwith's violet, Great Basin violet, sagebrush pansy, sagebrush violet |
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Habit | Plants perennial, caulescent, stoloniferous, 10–30 cm; stolons green or reddish, leafy, sometimes rooting at nodes, becoming lignified in age. | Plants perennial, caulescent, not stoloniferous, 2–22 cm. |
Stems | 1–5, prostrate, spreading, glabrous or sparsely puberulent, from current and/or previous year’s growth, on usually vertical, fleshy rhizome, rooting and forming rosettes at or near tip; rooted rosettes often develop into an erect, fleshy caudex from which new stems are produced. |
1–3, decumbent, ascending, or erect, ca. 1/2 subterranean, glabrous or usually puberulent, on single, short, vertical, deep-seated caudex. |
Leaves | evergreen, basal and cauline; basal: 1–6(–10); stipules deltate to ovate or linear-lanceolate, margins entire or glandular-toothed, apex acute to long-acuminate; petiole 2–16 cm, glabrous; blade often purple-spotted abaxially and/or adaxially, orbiculate to ovate, 1–4.5 × 2–3.9 cm, base cordate to truncate, margins crenate, eciliate, apex blunt to obtuse, mucronulate, surfaces glabrous or with scattered bristles on one or both surfaces; cauline similar to basal except: stipules deltate to lanceolate, margins entire or sparingly toothed; petiole 0.3–3 cm; blade 1.2–2.2 × 1.2–2 cm. |
basal and cauline; basal: 1–6 per caudex, palmately compound, ± 2-ternate or 3-ternate, leaflets 3; stipules adnate to petiole, forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute; petiole 2–10.5 cm, usually puberulent; blade ovate to deltate, 2.4–5 × 3.5–4.5 cm, base tapered, ultimate leaflets dissected into oblong, elliptic, lanceolate, or oblanceolate lobes 1–7 mm wide, lobe margins entire, usually ciliate, apex acute to obtuse, mucronulate, surfaces usually puberulent, seldom glabrous, abaxial surface without prominent vein parallel to each margin; cauline similar to basal except: stipules linear, apex acuminate; petiole 2–5.7 cm, usually puberulent, rarely glabrous; blade 1–2.7 × 1.5–3 cm. |
Peduncles | 5–10 cm, glabrous. |
1.5–10.6(–15.7) cm, usually puberulent, seldom glabrous. |
Flowers | sepals lanceolate to ovate-lanceolate, margins eciliate, auricles 1–2 mm; petals lemon-yellow on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded, lowest 8–17 mm, spur yellow or whitish, gibbous, 1–2.5 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins eciliate, auricles 0.1–1 mm; petals dark reddish violet on both surfaces, lower 3 usually lilac, rarely white or whitish, lateral 2 bearded, with yellow patch basally, dark reddish violet-veined, lowest 10–22 mm, with yellow patch, dark reddish violet-veined, spur whitish or yellowish, tinged purple, gibbous, 0.5–2 mm; style head bearded; cleistogamous flowers absent. |
Capsules | mottled with purple, spherical to ovoid, 5–8 mm, glabrous. |
oblong-ovoid, 7–12 mm, glabrous. |
Seeds | brown, tinged purple, 2–2.5 mm. |
brown, 3–4 mm. |
2n | = 24, 48. |
= 24. |
Viola sempervirens |
Viola beckwithii |
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Phenology | Flowering Jan–Jul. | Flowering Mar–May. |
Habitat | Redwood forests, other coastal forests, Douglas fir, other coniferous forests | Dry or moist places, among shrubs or beneath pines |
Elevation | 30–1400 m (100–4600 ft) | 900–2700 m (3000–8900 ft) |
Distribution |
AK; CA; ID; OR; WA; BC
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CA; ID; NV; OR; UT
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Discussion | In California, Viola sempervirens occurs in shaded redwood forests and other coastal forest habitats. In Oregon and Washington, it occurs in Douglas fir and other coniferous forests, where it can form mats (clones) one meter or more in diameter; its prostrate, spreading growth habit is similar to V. walteri. The leafy stems of V. sempervirens are similar to the leafy stolons of V. odorata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
In some populations of Viola beckwithii, the three lower petals are white with a yellow area proximally (V. B. Baird 1942). Leaves have been described as palmately biternate or triternate (L. Abrams and R. S. Ferris 1923–1960, vol. 3), ternately decompound into linear segments (C. L. Hitchcock et al. 1955–1969, vol. 3), palmately three-parted then bipinnately parted into ultimate linear or spatulate segments (P. A. Munz 1959), and palmately about three times three-parted into linear or spatulate-linear segments (W. L. Jepson 1951). Some populations in northern California are nearly or completely glabrous, which M. S. Baker recognized as var. glabrata. Viola beckwithii is reported to hybridize with V. utahensis (G. Davidse 1976). Observed pollinators of V. beckwithii in Utah include Apis mellifera Linnaeus and Anthophora ursina Cresson (Davidse). When Cottam described Viola bonnevillensis, he suggested that it could be a hybrid between V. beckwithii and V. utahensis, and G. Davidse (1976) concurred. The type specimen of V. bonnevillensis (Cottam 7067, UT) was examined by R. J. Little. Because no similar forms are known to have been collected since 1939, it is presumed that this taxon is a hybrid. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 157. | FNA vol. 6, p. 122. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. sarmentosa, V. sempervirens subsp. orbiculoides | V. beckwithii var. cachensis, V. beckwithii subsp. glabrata, V. bonnevillensis |
Name authority | Greene: Pittonia 4: 8. (1899) | Torrey: in E. M. Durand, Pl. Pratten. Calif., 82. (1855) |
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