Viola sempervirens |
Viola arvensis |
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evergreen violet, redwood violet, redwoods violet, trailing yellow violet, violette toujours verte |
European field violet, European field-pansy, field pansy, field violet, pansy, violette des champs, wild pansy |
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Habit | Plants perennial, caulescent, stoloniferous, 10–30 cm; stolons green or reddish, leafy, sometimes rooting at nodes, becoming lignified in age. | Plants annual, caulescent, not stoloniferous, 5–35 cm. |
Stems | 1–5, prostrate, spreading, glabrous or sparsely puberulent, from current and/or previous year’s growth, on usually vertical, fleshy rhizome, rooting and forming rosettes at or near tip; rooted rosettes often develop into an erect, fleshy caudex from which new stems are produced. |
1–5, usually erect, sometimes prostrate or decumbent, branched, subglabrous or puberulent, clustered on taproot. |
Leaves | evergreen, basal and cauline; basal: 1–6(–10); stipules deltate to ovate or linear-lanceolate, margins entire or glandular-toothed, apex acute to long-acuminate; petiole 2–16 cm, glabrous; blade often purple-spotted abaxially and/or adaxially, orbiculate to ovate, 1–4.5 × 2–3.9 cm, base cordate to truncate, margins crenate, eciliate, apex blunt to obtuse, mucronulate, surfaces glabrous or with scattered bristles on one or both surfaces; cauline similar to basal except: stipules deltate to lanceolate, margins entire or sparingly toothed; petiole 0.3–3 cm; blade 1.2–2.2 × 1.2–2 cm. |
cauline; stipules palmately lobed, middle lobe oblanceolate, obovate, elliptic, or lanceolate, ± equaling leaf blade, proximal lobes dissected, shorter, margins ciliate, apex acute to obtuse; petiole 0.5–2.3 cm, glabrous or puberulent; blade: proximal ovate to ± oblong, distal narrowly or broadly lanceolate, 0.8–3.4 × 0.3–1.9 cm, base attenuate to ± truncate, margins coarsely crenate-serrate, ciliate or eciliate, apex acute to obtuse, surfaces pubescent abaxially, at least on major veins, glabrous adaxially. |
Peduncles | 5–10 cm, glabrous. |
2–8 cm, glabrous or pubescent. |
Flowers | sepals lanceolate to ovate-lanceolate, margins eciliate, auricles 1–2 mm; petals lemon-yellow on both surfaces, lower 3 and sometimes upper 2 brownish purple-veined, lateral 2 bearded, lowest 8–17 mm, spur yellow or whitish, gibbous, 1–2.5 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate, margins ciliate or eciliate, auricles 2–4 mm; petals white to pale yellow on both surfaces, upper 2 ± violet, lower 3 with yellow basal area, often violet-veined, lateral 2 bearded, ± equaling or shorter than sepals, lowest with dark yellow area basally, 7–15 mm, spur blue-violet to purple, elongated, 3–5 mm; style head bearded; cleistogamous flowers absent. |
Capsules | mottled with purple, spherical to ovoid, 5–8 mm, glabrous. |
± spherical, 5–9 mm, glabrous. |
Seeds | brown, tinged purple, 2–2.5 mm. |
brown, 1.5–1.9 mm. |
2n | = 24, 48. |
= 34. |
Viola sempervirens |
Viola arvensis |
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Phenology | Flowering Jan–Jul. | Flowering May–Jul. |
Habitat | Redwood forests, other coastal forests, Douglas fir, other coniferous forests | Abandoned fields, roadsides, lawns |
Elevation | 30–1400 m (100–4600 ft) | 0–3000 m (0–9800 ft) |
Distribution |
AK; CA; ID; OR; WA; BC
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CA; CT; DC; DE; GA; ID; IL; IN; KS; KY; LA; MA; MD; ME; MI; MN; MO; MS; MT; NC; NE; NH; NJ; NY; OH; OR; PA; RI; SC; TN; UT; VA; VT; WA; WI; WV; AB; BC; MB; NB; NL; NS; ON; PE; QC; SK; SPM; Asia (Siberia); Greenland; Europe; Africa [Introduced in North America]
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Discussion | In California, Viola sempervirens occurs in shaded redwood forests and other coastal forest habitats. In Oregon and Washington, it occurs in Douglas fir and other coniferous forests, where it can form mats (clones) one meter or more in diameter; its prostrate, spreading growth habit is similar to V. walteri. The leafy stems of V. sempervirens are similar to the leafy stolons of V. odorata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
A. R. Clapham et al. (1987) noted that Viola arvensis is pollinated by insects and is often selfed; T. Marcussen and T. Karlsson (2010) stated that V. arvensis regularly self-pollinates. Roots of V. arvensis have the odor of wintergreen when crushed (A. E. Radford et al. 1968; W. J. Hayden and J. Clough 1990). Viola arvensis is not vegetatively distinguishable from V. tricolor var. tricolor. G. Halliday (pers. comm.) reported that Viola arvensis occurs in southwestern Greenland, where it is introduced, and is ephemeral on St. Pierre and Miquelon. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 157. | FNA vol. 6, p. 121. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | V. sarmentosa, V. sempervirens subsp. orbiculoides | Mnemion arvense, V. tricolor var. arvensis |
Name authority | Greene: Pittonia 4: 8. (1899) | Murray: Prodr. Stirp. Gott., 73. (1770) |
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