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great-spur or long-spur or Selkirk's violet, great-spur violet, Selkirk's violet, violette de Selkirk

Utah violet

Habit Plants perennial, acaulescent, not stoloniferous, 4–15 cm; rhizome slender, not fleshy. Plants perennial, caulescent, not stoloniferous, 5–20 cm.
Stems

1–4, prostrate, decumbent, or erect, leafy proximally and distally, glabrate or sparsely puberulent, on caudex from subligneous rhizome.

Leaves

basal, 2–12, prostrate to ascending;

stipules linear-lanceolate, margins entire, apex acute;

petiole 1.5–7 cm, not winged, glabrous or pubescent;

blade unlobed, usually ovate, rarely orbiculate, 1–5 × 1–5 cm, base cordate, margins crenate to crenulate or serrate, eciliate, apex rounded to acute, surfaces glabrous or sometimes pubescent abaxially, strigose adaxially.

basal and cauline;

basal: 1–3;

stipules adnate to petiole, forming 2 linear-lanceolate wings, margins entire or laciniate with glandular hairs, apex of each wing free, acute, sometimes divided into several filiform processes;

petiole 2–8(–12) cm, glabrate or sparsely puberulent;

blade usually broadly ovate, sometimes ± orbiculate, 1–4.5 × 0.7–3.4 cm, base usually attenuate, rarely truncate or subcordate, margins ± coarsely crenate-serrate, ciliate, apex obtuse, surfaces glabrate or sparsely puberulent;

cauline similar to basal except: stipules lanceolate or linear-lanceolate to linear, margins ± entire or lacerate to laciniate with sparse gland-tipped projections, apex acute to acuminate, often divided into 2–4 filiform processes;

petiole 3–8.7 cm;

blade ovate to oblong-ovate or oblong-lanceolate, 0.7–1.6 × 2.6–3.7 cm, margins shallowly serrate, irregularly crenate, or ± entire, apex acute to obtuse.

Peduncles

3–6 cm, glabrous or pubescent.

4–10 cm, glabrate or sparsely puberulent.

Flowers

sepals lanceolate to ovate-lanceolate, margins eciliate, auricles 1–2 mm;

petals light violet on both surfaces, lower 3 white basally and dark violet-veined, lateral 2 beardless, lowest 8–13 mm, spur pale to dark violet, elongated, 4–7 mm;

style head beardless; cleistogamous flowers on prostrate to ascending peduncles.

sepals lanceolate, margins ciliate or eciliate, auricles 1–1.5 mm;

petals deep lemon-yellow adaxially, upper 2 and sometimes lateral 2 reddish brown to brownish purple abaxially, lower 3 and sometimes upper 2 dark brown-veined, lateral 2 sparsely bearded, lowest 10–13 mm, spur yellow, gibbous, 0.5–1.5 mm;

style head bearded; cleistogamous flowers axillary.

Capsules

ovoid to ellipsoid, 4–8 mm, glabrous.

± spherical, 6–7 mm, glabrous or finely puberulent.

Seeds

brown, 1–2 mm.

brown, ca. 3 mm.

2n

= 24.

= 24.

Viola selkirkii

Viola utahensis

Phenology Flowering May–Jun. Flowering May.
Habitat Wet to moist places, thickets, mixed or coniferous woods Open sagebrush areas, semidesert slopes, grasslands, open forests
Elevation 200–3000 m (700–9800 ft) 1200–2600 m (3900–8500 ft)
Distribution
from FNA
AK; CO; MA; ME; MI; MN; NH; NY; PA; SD; VT; WA; WI; AB; BC; MB; NB; NL; NS; ON; QC; SK; YT; Greenland; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
CO; ID; NV; UT; WY
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola selkirkii occurs on the southwestern coast of Greenland, north to 63ºN (L. Brouillet, pers. comm.). Presence of V. selkirkii in the Northwest Territories and Nunavut is considered doubtful (L. Brouillet et al., http://canadensys.net/vascan). K. W. Allred (2008) said that V. selkirkii was considered by W. C. Martin and C. R. Hutchins (1980) to be expected in New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola utahensis occurs only in areas where its closest relatives, V. vallicola and V. purpurea var. venosa, are also found and it appears to have originated from the crossing of those taxa followed by a doubling of the chromosomes (M. S. Baker 1949). The elaiosome of V. utahensis seeds has the same general appearance as those of other members of V. purpurea complex (except V. purpurea var. venosa), covering about one-third the length of the seed. G. Davidse (1976) found that tetraploid members of Clausen’s “subsect.” Purpurea (V. utahensis) have serrate leaf margins; diploid plants (V. purpurea) have crenate leaf margins. He also found that the size of V. utahensis plants is correlated with elevation, the largest plants at low elevations and the smallest at higher elevations.

G. Davidse (1976) reported putative hybrids between Viola utahensis and V. praemorsa subsp. major (= var. linguifolia), V. purpurea var. venosa, and V. beckwithii.

Observed pollinators of Viola utahensis in Utah include the bees Apis mellifera Linnaeus and Anthophora ursina Cresson (G. Davidse 1976). In other populations, flies in the genera Eristalis Latreille and Bombylius Linnaeus were observed indiscriminately visiting the yellow-flowered species V. praemorsa subsp. major, V. utahensis, and V. vallicola (Davidse).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 157. FNA vol. 6, p. 162.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. vallicola, V. villosa, V. walteri
Name authority Pursh ex Goldie: Edinburgh Philos. J. 6: 324. (1822) M. S. Baker & J. C. Clausen: Leafl. W. Bot. 5: 145. (1949)
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