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great-spur or long-spur or Selkirk's violet, great-spur violet, Selkirk's violet, violette de Selkirk

early blue violet, three-lobe violet, three-lobed violet, trilobed violet, wood violet

Habit Plants perennial, acaulescent, not stoloniferous, 4–15 cm; rhizome slender, not fleshy. Plants perennial, acaulescent, not stoloniferous, 6–50 cm; rhizome thick, fleshy.
Leaves

basal, 2–12, prostrate to ascending;

stipules linear-lanceolate, margins entire, apex acute;

petiole 1.5–7 cm, not winged, glabrous or pubescent;

blade unlobed, usually ovate, rarely orbiculate, 1–5 × 1–5 cm, base cordate, margins crenate to crenulate or serrate, eciliate, apex rounded to acute, surfaces glabrous or sometimes pubescent abaxially, strigose adaxially.

basal, 2–3, ascending to erect;

stipules linear-lanceolate, margins entire, apex acute;

petiole 1–20 cm, glabrous or pubescent;

earliest leaf blades unlobed, mid-season blades 3–9-lobed, lobes sometimes appearing petiolate and sometimes further lobed;

earliest leaf blades reniform to ovate, mid-season blades with middle lobes usually ovate or elliptic to widely obovate, sometimes narrowly elliptic, narrowly ovate, lanceolate, or spatulate to narrowly obovate, lateral lobes elliptic, obdeltate, or spatulate to falcate, smaller lobes similar, 1–14 × 1–10 cm, base truncate to cordate, margins serrate or entire, usually ciliate, apex acute, rounded, blunt, or obtuse, surfaces glabrous or pubescent throughout or along veins.

Peduncles

3–6 cm, glabrous or pubescent.

3–13 cm, glabrous or pubescent.

Flowers

sepals lanceolate to ovate-lanceolate, margins eciliate, auricles 1–2 mm;

petals light violet on both surfaces, lower 3 white basally and dark violet-veined, lateral 2 beardless, lowest 8–13 mm, spur pale to dark violet, elongated, 4–7 mm;

style head beardless; cleistogamous flowers on prostrate to ascending peduncles.

sepals lanceolate, margins ciliate or eciliate, auricles 0.5–1 mm;

petals violet on both surfaces, lower 3 white basally, lower 3 and upper 2 sometimes purple-veined, lateral 2 bearded, spur sometimes bearded, lowest 15–25 mm, spur white, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers on prostrate to ascending peduncles.

Capsules

ovoid to ellipsoid, 4–8 mm, glabrous.

ellipsoid, 5–15 mm, glabrous.

Seeds

brown, 1–2 mm.

beige, mottled to bronze, 1.5–2.5 mm.

2n

= 24.

= 54.

Viola selkirkii

Viola palmata

Phenology Flowering May–Jun.
Habitat Wet to moist places, thickets, mixed or coniferous woods
Elevation 200–3000 m (700–9800 ft)
Distribution
from FNA
AK; CO; MA; ME; MI; MN; NH; NY; PA; SD; VT; WA; WI; AB; BC; MB; NB; NL; NS; ON; QC; SK; YT; Greenland; Eurasia
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; CT; DC; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; ME; MI; MO; MS; NC; ND; NE; NJ; NM; OH; OK; PA; RI; SC; TN; TX; VA; VT; WV; ON
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola selkirkii occurs on the southwestern coast of Greenland, north to 63ºN (L. Brouillet, pers. comm.). Presence of V. selkirkii in the Northwest Territories and Nunavut is considered doubtful (L. Brouillet et al., http://canadensys.net/vascan). K. W. Allred (2008) said that V. selkirkii was considered by W. C. Martin and C. R. Hutchins (1980) to be expected in New Mexico.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

N. L. Gil-Ad (1997) presented evidence based on seed coat micromorphology that some plants known as Viola palmata are hybrids between plants with lobed and unlobed leaves and believed the type specimen to be hybrid in origin. He chose not to recognize the name V. palmata, choosing instead to recognize V. triloba. If the type specimen does represent a hybrid, Gil-Ad would be correct in recognizing V. triloba. Because the purity of the type specimen cannot be ascertained, we consider V. palmata the most appropriate name for this taxon. We acknowledge the presence of hybrids between lobed and unlobed plants; such hybrids obscure lines of demarcation. Some herbarium specimens may represent such hybrids; we feel most do not.

L. E. McKinney (1992) described the nomenclatural history of Viola palmata. Homophylly versus heterophylly differentiates V. palmata from V. subsinuata, the homophyllous violet most often called V. palmata. We know that the taxon described by Linnaeus was heterophyllous, with some undivided leaf blades.

Varieties 2 (2 in the flora).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Petioles, leaf surfaces, and peduncles usually pubescent, rarely glabrous; apex of middle leaf lobe acute; plants of dry to mesic habitats.
var. palmata
1. Petioles, leaf surfaces, and peduncles usually glabrous, rarely pubescent; apex of middle leaf lobe rounded, blunt, or obtuse; plants of wet ground.
var. heterophylla
Source FNA vol. 6, p. 157. FNA vol. 6, p. 141.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Subordinate taxa
V. palmata var. heterophylla, V. palmata var. palmata
Synonyms V. cucullata var. palmata
Name authority Pursh ex Goldie: Edinburgh Philos. J. 6: 324. (1822) Linnaeus: Sp. Pl. 2: 933. (1753)
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