Viola rostrata |
Viola subsinuata |
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long-spur violet, violette rostrée |
early blue violet, wavy-leaf violet, wood violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 5–20 cm. | Plants perennial, acaulescent, not stoloniferous, 10–30 cm; rhizome thick, fleshy. |
Stems | 1–7, ascending to erect (often declining during anthesis), glabrous, on caudex from fleshy rhizome. |
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Leaves | basal and cauline; basal: 1–5; stipules lanceolate, margins laciniate, apex acute; petiole 1–9.6 cm, glabrous; blade sometimes purple-spotted abaxially and/or adaxially, reniform to ovate, 1–4.5 × 1–4 cm, base broadly cordate to cordate, margins crenate to serrate, mostly eciliate, apex obtuse to acute, surfaces usually pubescent, mostly adaxially toward base, sometimes glabrous; cauline similar to basal except: petiole 0.4–4 cm; distal cauline blades ovate to deltate, 1–4.5 × 1–4 cm, base cordate, apex acuminate to acute. |
basal, 2–11, ascending to erect; stipules linear-lanceolate, margins entire, apex acute; petiole 5–25 cm, glabrous or pubescent; mid-season leaf blades incised or lobed throughout, earliest leaf blades lobed (plants homophyllous), similar to mid-season blades, blade 5–9(–16)-lobed, sinuses usually narrower, shallower toward leaf base, middle and lateral blade lobes differ in width and/or shape, middle lobes narrowly deltate to narrowly elliptic, lateral lobes narrowly elliptic, lanceolate, or falcate, 1–11 × 1–12 cm, base truncate to cordate, margins entire or crenate, ciliate or eciliate, apex acute, mucronulate, surfaces usually pubescent throughout or on veins. |
Peduncles | 5–9 cm, usually glabrous. |
5–15 cm, glabrous or pubescent. |
Flowers | sepals lanceolate, margins eciliate, auricles 1–2 mm; petals pale lavender-violet on both surfaces, all white basally, lower 3 purple-black-veined, all beardless, lowest 8–20 mm, spur white, purple, or lavender-tinged, elongated, 10–20 mm; style head beardless; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to dark blue-violet on both surfaces, lower 3 and upper 2 sometimes purple-veined, lateral 2 bearded, lowest sometimes bearded, 15–25 mm, spur color same as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
Capsules | ellipsoid, 4–6 mm, glabrous. |
ellipsoid, 8–12 mm, glabrous. |
Seeds | beige to bronze, 1.3–2 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
2n | = 20. |
= 54. |
Viola rostrata |
Viola subsinuata |
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Phenology | Flowering Apr–May. | Flowering Apr–Jun. |
Habitat | Rich, mesic to dry, well-drained woodlands, mountains | Rich woods |
Elevation | 200–1800 m (700–5900 ft) | 100–3000 m (300–9800 ft) |
Distribution |
AL; CT; GA; IN; KY; MA; MD; MI; NC; NJ; NY; OH; PA; SC; TN; VA; VT; WI; WV; ON; QC; e Asia (Japan)
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CT; DC; DE; GA; IL; KY; MA; MD; MI; NC; NJ; NY; OH; PA; RI; SC; TN; VA; VT; WI; WV; ON
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Discussion | Viola rostrata has the longest spur of any North American Viola species. Viola rostrata reportedly hybridizes with V. labradorica (= V. ×malteana House) and V. striata (= V. ×brauniae Grover ex Cooperrider). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
L. E. McKinney (1992) described the misconceptions surrounding Viola subsinuata that many came to think of as the heterophyllous V. palmata, whereas V. subsinuata is homophyllous. As E. Brainerd (1910, 1921) pointed out, heterophylly versus homophylly is an important and steadfast character difference in acaulescent Viola species. N. L. Gil-Ad (1997) chose not to recognize V. subsinuata, suggesting that it represented a hybrid or introgressant. H. E. Ballard (2000) and A. Haines (2011) both recognized V. subsinuata as a species. Viola subsinuata is a state historical species in Rhode Island, where it was last documented in 1941 (R. W. Enser, http://rinhs.org/wp-content/uploads/ri_rare_plants_2007.pdf). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 154. | FNA vol. 6, p. 159. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | Lophion rostratum | V. emarginata var. subsinuata, V. palmata var. angelliae |
Name authority | Pursh: Fl. Amer. Sept. 1: 174. (1813) | (Greene) Greene: Pittonia 4: 4. (1899) |
Web links |