Viola rostrata |
Viola guadalupensis |
|
---|---|---|
long-spur violet, violette rostrée |
Guadalupe Mountains violet, Guadalupe violet |
|
Habit | Plants perennial, caulescent, not stoloniferous, 5–20 cm. | Plants perennial, caulescent, not stoloniferous, 1–10 cm. |
Stems | 1–7, ascending to erect (often declining during anthesis), glabrous, on caudex from fleshy rhizome. |
1–5, decumbent to erect, leafy proximally and distally, glabrous, on caudex from fleshy rhizome. |
Leaves | basal and cauline; basal: 1–5; stipules lanceolate, margins laciniate, apex acute; petiole 1–9.6 cm, glabrous; blade sometimes purple-spotted abaxially and/or adaxially, reniform to ovate, 1–4.5 × 1–4 cm, base broadly cordate to cordate, margins crenate to serrate, mostly eciliate, apex obtuse to acute, surfaces usually pubescent, mostly adaxially toward base, sometimes glabrous; cauline similar to basal except: petiole 0.4–4 cm; distal cauline blades ovate to deltate, 1–4.5 × 1–4 cm, base cordate, apex acuminate to acute. |
cauline; stipules lanceolate to ovate- or oblong-lanceolate or linear, margins sparingly glandular-fimbriate, apex acute; petiole 2–6 cm, glabrous; blade ovate to ovate-deltate or ovate-lanceolate, 1.2–2.4 × 0.7–1.3 cm, base broadly cuneate to rounded or truncate, margins entire or with 1–3 crenations on proximal 1/2, eciliate, apex acute to rounded, surfaces glabrous, sometimes with a few short hairs on veins abaxially. |
Peduncles | 5–9 cm, usually glabrous. |
3.5–6 cm, glabrous. |
Flowers | sepals lanceolate, margins eciliate, auricles 1–2 mm; petals pale lavender-violet on both surfaces, all white basally, lower 3 purple-black-veined, all beardless, lowest 8–20 mm, spur white, purple, or lavender-tinged, elongated, 10–20 mm; style head beardless; cleistogamous flowers axillary. |
sepals linear to linear-lanceolate, margins eciliate, auricles 0.5–1.5 mm; petals deep lemon-yellow adaxially, upper 2 reddish brown abaxially, lateral 2 and lowest dark brown-veined basally, lateral 2 bearded, lowest 7–11 mm, spur yellow, gibbous, 1–1.4 mm; style head bearded; cleistogamous flowers absent. |
Capsules | ellipsoid, 4–6 mm, glabrous. |
ovoid, 3–4.5 mm, glabrous. |
Seeds | beige to bronze, 1.3–2 mm. |
light brown, ± 2 mm. |
2n | = 20. |
= 24. |
Viola rostrata |
Viola guadalupensis |
|
Phenology | Flowering Apr–May. | Flowering May. |
Habitat | Rich, mesic to dry, well-drained woodlands, mountains | Openings and narrow ledges on limestone rock faces |
Elevation | 200–1800 m (700–5900 ft) | 2600 m (8500 ft) |
Distribution |
AL; CT; GA; IN; KY; MA; MD; MI; NC; NJ; NY; OH; PA; SC; TN; VA; VT; WI; WV; ON; QC; e Asia (Japan)
|
TX |
Discussion | Viola rostrata has the longest spur of any North American Viola species. Viola rostrata reportedly hybridizes with V. labradorica (= V. ×malteana House) and V. striata (= V. ×brauniae Grover ex Cooperrider). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Viola guadalupensis is known only from the eastern rim of the Guadalupe Mountains in Culberson County. Powell and Wauer noted that it is the only yellow-flowered violet known in the Guadalupe Mountains and appears to be related to V. nuttallii and V. vallicola. K. W. Allred (2008) stated that a report of this species in New Mexico by J. T. Kartesz and C. A. Meacham (1999) requires verification. K. Haskins (pers. comm.) reported that experiments are currently being conducted to propagate plants of V. guadalupensis via cell tissue culture. Chloroplast (trnL-F spacer) and low-copy nuclear gene (GPI) phylogenies indicate that Viola guadalupensis is an alloploid that originated through hybridization between an unidentified member of subsect. Canadenses (the paternal parent) and a member of the V. nuttallii complex (the maternal parent), of sect. Chamaemelanium (T. Marcussen et al. 2011). Evidence reported by these authors from a fossil-calibrated relaxed clock dating analysis showed the estimated maximum age of V. guadalupensis to be (5.7–)8.6(–11.6) million years. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 6, p. 154. | FNA vol. 6, p. 132. |
Parent taxa | Violaceae > Viola | Violaceae > Viola |
Sibling taxa | ||
Synonyms | Lophion rostratum | |
Name authority | Pursh: Fl. Amer. Sept. 1: 174. (1813) | A. M. Powell & Wauer: Sida 14: 1, fig. 1. (1990) |
Web links |