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long-spur violet, violette rostrée

Douglas' golden violet, Douglas' or Douglas' golden violet, Douglas' violet, Douglas' yellow violet, golden violet

Habit Plants perennial, caulescent, not stoloniferous, 5–20 cm. Plants perennial, caulescent, not stoloniferous, 3–20 cm.
Stems

1–7, ascending to erect (often declining during anthesis), glabrous, on caudex from fleshy rhizome.

1–3, decumbent or ascending to erect, ca. 1/2 subterranean, glabrous or puberulent, from single, short, vertical, deep-seated caudex.

Leaves

basal and cauline;

basal: 1–5;

stipules lanceolate, margins laciniate, apex acute;

petiole 1–9.6 cm, glabrous;

blade sometimes purple-spotted abaxially and/or adaxially, reniform to ovate, 1–4.5 × 1–4 cm, base broadly cordate to cordate, margins crenate to serrate, mostly eciliate, apex obtuse to acute, surfaces usually pubescent, mostly adaxially toward base, sometimes glabrous;

cauline similar to basal except: petiole 0.4–4 cm;

distal cauline blades ovate to deltate, 1–4.5 × 1–4 cm, base cordate, apex acuminate to acute.

basal and cauline;

basal: 1–6, bipinnately compound, leaflets 3–5;

stipules adnate to petiole forming 2 linear-lanceolate wings, unlobed, margins entire, apex of each wing free, acute to acuminate;

petiole 5–6.8 cm, glabrous or puberulent;

blade ovate, 3.5–5 × 2.4–3.5 cm, base tapered, leaflets 3–5-lobed, lobes linear, narrowly elliptic, or oblong, 1–2.5(–5) mm wide, margins entire, usually densely ciliate, apex acute to obtuse, mucronulate, surfaces glabrous or puberulent;

cauline similar to basal except: stipules ovate to linear-lanceolate, margins entire or toothed, apex acute;

petiole 0.9–4 cm;

blade 1.1–4.1 × 1–3.6 cm.

Peduncles

5–9 cm, usually glabrous.

2–12.5 cm, glabrous or puberulent.

Flowers

sepals lanceolate, margins eciliate, auricles 1–2 mm;

petals pale lavender-violet on both surfaces, all white basally, lower 3 purple-black-veined, all beardless, lowest 8–20 mm, spur white, purple, or lavender-tinged, elongated, 10–20 mm;

style head beardless; cleistogamous flowers axillary.

sepals lanceolate, margins ciliate, auricles 0.5–1.5 mm;

petals light golden yellow adaxially, upper 2 dark brown to ± black abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 8–21 mm, spur dark greenish to dark brown, gibbous, 1.5–2 mm;

style head bearded; cleistogamous flowers absent.

Capsules

ellipsoid, 4–6 mm, glabrous.

spherical to oblong, 5–12 mm, glabrous.

Seeds

beige to bronze, 1.3–2 mm.

light brown, 2.8–3.3 mm.

2n

= 20.

= 24, 48.

Viola rostrata

Viola douglasii

Phenology Flowering Apr–May. Flowering Feb–Jul.
Habitat Rich, mesic to dry, well-drained woodlands, mountains Vernally moist grassy slopes and flats, often serpentine soil (except Oregon)
Elevation 200–1800 m (700–5900 ft) 20–2300 m (100–7500 ft)
Distribution
from FNA
AL; CT; GA; IN; KY; MA; MD; MI; NC; NJ; NY; OH; PA; SC; TN; VA; VT; WI; WV; ON; QC; e Asia (Japan)
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; Mexico (Baja California)
[WildflowerSearch map]
[BONAP county map]
Discussion

Viola rostrata has the longest spur of any North American Viola species.

Viola rostrata reportedly hybridizes with V. labradorica (= V. ×malteana House) and V. striata (= V. ×brauniae Grover ex Cooperrider).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola douglasii is tetraploid (n = 12) south of, and octoploid (n = 24) north of, San Francisco Bay, California. It forms sterile hybrids with V. quercetorum (J. Clausen 1964). V. B. Baird (1936) described V. douglasii × purpurea, which Clausen later said was actually V. quercetorum, not described at the time of Baird’s publication.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 154. FNA vol. 6, p. 129.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms Lophion rostratum V. chrysantha
Name authority Pursh: Fl. Amer. Sept. 1: 174. (1813) Steudel: Nomencl. Bot. ed. 2, 2. 771. (1841)
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