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kidney-leaf or kidney-shape or white violet, kidney-leaf violet, kidney-leaf white violet, northern white violet, violette réniforme, white violet

Howell's violet, violette de Howell

Habit Plants perennial, acaulescent, not stoloniferous, 5–30 cm; rhizome thick, fleshy. Plants perennial, caulescent, not stoloniferous, 2–44 cm.
Stems

1–4, ascending to erect (sometimes later reclining to nearly prostrate), glabrous or sparsely pubescent, on caudex from subligneous rhizome.

Leaves

basal, 1–5, prostrate to ascending;

stipules linear-lanceolate, margins entire to sparsely laciniate, apex acute;

petiole 3–10 cm, strigose, sericeous, or villous, occasionally glabrous;

blade unlobed, reniform or ovate to broadly ovate or orbiculate, 1.5–3.5 × 2–5 cm, base cordate to broadly cordate, margins serrate-crenate, ciliate or eciliate, apex acute, obtuse, or rounded, occasionally acuminate, surfaces usually sparsely to densely strigose, sericeous, or villous throughout or along veins, sometimes glabrous.

basal and cauline;

basal: 1–6;

stipules linear-lanceolate to oblong, margins mostly entire or laciniate with gland-tipped projections, apex acute to acuminate;

petiole 4–15 cm, glabrous or sparsely puberulent;

blade ovate to reniform, 1.9–6.8 × 2.1–6.4 cm, thin, base cordate, margins crenate, ciliate, apex acute to usually obtuse, surfaces glabrous or sparsely puberulent;

cauline similar to basal except: stipules linear-lanceolate or deltate to oblong, apex acute or long-acuminate;

petiole 1–4.5 cm;

blade 2–5.1 × 1.2–5.8 cm.

Peduncles

3–8 cm, puberulent.

2.8–17.8 cm, glabrous or sparsely pubescent.

Flowers

sepals lanceolate to ovate, margins usually eciliate, auricles 1–2 mm;

petals white on both surfaces, lower 3 purple-veined, all beardless or lower 3 sparsely bearded, lowest 8–10 mm, spur white, gibbous, 2–3 mm;

style head beardless; cleistogamous flowers on prostrate to ascending peduncles.

sepals lanceolate, margins ciliate or eciliate, auricles 1–2 mm;

petals violet to soft blue-violet on both surfaces, whitish basally, lower 3 dark violet-veined, lateral 2 densely bearded, lowest 14–23 mm, spur blue-violet to whitish, gibbous to elongated, 2.4–5 mm, usually less than 1/2 lowest petal, tip straight;

style head bearded; cleistogamous flowers axillary.

Capsules

ovoid to ellipsoid, 5–8 mm, glabrous.

ellipsoid, 7–11 mm, glabrous.

Seeds

mottled beige to bronze, 1.5–2.2 mm.

light brown, 2.1–2.8 mm.

2n

= 24.

= 40, 80.

Viola renifolia

Viola howellii

Phenology Flowering Apr–Jun. Flowering Apr–Jul.
Habitat Moist, often shaded alluvial or upland forests, shrub thickets, stream banks, swamp forests, bogs, fens Moist, shady areas, coniferous forests, prairies, along streams
Elevation 200–3000 m (700–9800 ft) 50–1500 m (200–4900 ft)
Distribution
from FNA
AK; CO; CT; IA; ID; MA; ME; MI; MN; MT; NH; NY; PA; SD; VT; WA; WI; WY; AB; BC; MB; NB; NL; NS; NT; NU; ON; PE; QC; SK; YT
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; OR; WA; BC
[WildflowerSearch map]
[BONAP county map]
Discussion

Nonflowering plants of Viola renifolia and V. epipsila appear similar. The abaxial leaf surfaces of V. renifolia usually have a few short, straight hairs on the main veins; leaves of V. epipsila are usually glabrous (C. Parker, pers. comm.). Viola renifolia can appear similar to V. macloskeyi. V. B. Baird (1942) wrote that V. renifolia sometimes produces ascending stems. H. A. Gleason and A. Cronquist (1991) noted that if stolons were present, they were short and racemelike with cleistogamous flowers. The ascending stems and stolons mentioned by these authors may or may not be different phases of the same structure.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Viola howellii is similar to V. adunca and usually occupies moister habitats along the western side of the Cascade Mountains (H. E. Ballard 1992). The first documented record of this species in California was recently reported from Siskiyou County (R. J. Little 2010).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 6, p. 153. FNA vol. 6, p. 134.
Parent taxa Violaceae > Viola Violaceae > Viola
Sibling taxa
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. howellii, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
V. adunca, V. affinis, V. arvensis, V. bakeri, V. beckwithii, V. bicolor, V. biflora, V. blanda, V. brittoniana, V. canadensis, V. canina, V. charlestonensis, V. clauseniana, V. cucullata, V. cuneata, V. douglasii, V. egglestonii, V. epipsila, V. flettii, V. frank-smithii, V. glabella, V. guadalupensis, V. hallii, V. hastata, V. hirsutula, V. japonica, V. labradorica, V. lanceolata, V. langsdorffii, V. lithion, V. lobata, V. macloskeyi, V. missouriensis, V. nephrophylla, V. novae-angliae, V. nuttallii, V. ocellata, V. odorata, V. orbiculata, V. palmata, V. palustris, V. pedata, V. pedatifida, V. pedunculata, V. pinetorum, V. praemorsa, V. primulifolia, V. prionantha, V. pubescens, V. purpurea, V. quercetorum, V. renifolia, V. riviniana, V. rostrata, V. rotundifolia, V. sagittata, V. selkirkii, V. sempervirens, V. septemloba, V. sheltonii, V. sororia, V. striata, V. subsinuata, V. tomentosa, V. tricolor, V. trinervata, V. tripartita, V. umbraticola, V. utahensis, V. vallicola, V. villosa, V. walteri
Synonyms V. blanda var. renifolia, V. brainerdii, V. mistassinica, V. renifolia var. brainerdii
Name authority A. Gray: Proc. Amer. Acad. Arts 8: 288. (1870) A. Gray: Proc. Amer. Acad. Arts 22: 308. (1887)
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