Viola renifolia
Viola brittoniana
kidney-leaf or kidney-shape or white violet, kidney-leaf violet, kidney-leaf white violet, northern white violet, violette réniforme, white violet
coast violet, coastal violet, northern coastal violet
basal, 1–5, prostrate to ascending;
stipules linear-lanceolate, margins entire to sparsely laciniate, apex acute;
petiole 3–10 cm, strigose, sericeous, or villous, occasionally glabrous;
blade unlobed, reniform or ovate to broadly ovate or orbiculate, 1.5–3.5 × 2–5 cm, base cordate to broadly cordate, margins serrate-crenate, ciliate or eciliate, apex acute, obtuse, or rounded, occasionally acuminate, surfaces usually sparsely to densely strigose, sericeous, or villous throughout or along veins, sometimes glabrous.
basal, 5–9, ascending to erect, 5–9-lobed;
stipules linear-lanceolate, margins entire, apex obtuse;
petiole 3–16 cm, usually glabrous;
mid-season blades incised or lobed throughout, earliest blades lobed (plants homophyllous), similar to mid-season blades, middle and lateral blade lobes differ in width and/or shape, middle lobes lanceolate or spatulate to narrowly obovate, lateral lobes lanceolate or spatulate to falcate (each sometimes with deltate to falcate appendages or teeth along margins), 1–7 × 2–8 cm, base truncate to cordate, margins entire, ciliate, apex acute to obtuse, surfaces usually glabrous, rarely with a few strigose, hairs concentrated on veins.
3–8 cm, puberulent.
5–18 cm, usually glabrous.
sepals lanceolate to ovate, margins usually eciliate, auricles 1–2 mm;
petals white on both surfaces, lower 3 purple-veined, all beardless or lower 3 sparsely bearded, lowest 8–10 mm, spur white, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on prostrate to ascending peduncles.
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 2–3 mm;
petals light to soft reddish violet on both surfaces, lower 3 white basally, dark violet-veined, lateral 2 bearded, lowest 10–25 mm, sometimes bearded, spur same color as petals, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on ascending to erect peduncles.
ovoid to ellipsoid, 5–8 mm, glabrous.
ellipsoid, 10–15 mm, glabrous.
mottled beige to bronze, 1.5–2.2 mm.
beige, mottled to bronze, 1.5–2.5 mm.
= 24.
= 54.
Viola renifolia
Viola brittoniana
Nonflowering plants of Viola renifolia and V. epipsila appear similar. The abaxial leaf surfaces of V. renifolia usually have a few short, straight hairs on the main veins; leaves of V. epipsila are usually glabrous (C. Parker, pers. comm.). Viola renifolia can appear similar to V. macloskeyi. V. B. Baird (1942) wrote that V. renifolia sometimes produces ascending stems. H. A. Gleason and A. Cronquist (1991) noted that if stolons were present, they were short and racemelike with cleistogamous flowers. The ascending stems and stolons mentioned by these authors may or may not be different phases of the same structure.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Britton considered Viola brittoniana similar to V. pedatifida, apparently making him the first to recognize their affinity. Both are homophyllous and similar in other characters (L. E. McKinney 1992). McKinney recognized V. brittoniana as a variety of V. pedatifida. Additional evidence (N. L. Gil-Ad 1997; A. Haines 2011) suggests that these taxa are sufficiently distinct to continue recognition as separate species.
Viola pectinata has sharply dentate leaves and is closely related to V. brittoniana, usually occurring with it. N. L. Gil-Ad (1997) made a convincing argument to recognize it as a form; A. Haines (2011) recognized it as a species. Others have considered it to be a sporadic form that may be of hybrid origin, or, as N. H. Russell (1965) suggested, is due to genetic dimorphism. We are reserving a decision on treating this taxon until additional studies are completed.
A. Haines (2011) reported that the distribution of Viola brittoniana in Maine is actually based on the hybrid V. ×insolita House (V. brittoniana × V. sororia).
Viola brittoniana reportedly hybridizes with V. cucullata (= V. ×notabilis E. P. Bicknell), V. sagittata var. sagittata (= V. ×mulfordiae Pollard), and V. sororia (= V. ×insolita).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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