Viola purpurea |
Viola guadalupensis |
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goosefoot violet, mountain violet, pine violet, purple-mark yellow violet |
Guadalupe Mountains violet, Guadalupe violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 1.5–25 cm. | Plants perennial, caulescent, not stoloniferous, 1–10 cm. | ||||||||||||||||||||||||
Stems | 1–5(–7), decumbent or spreading to erect, leafy proximally and distally, ± glabrous, puberulent, canescent, or tomentose, on caudex from subligneous rhizome. |
1–5, decumbent to erect, leafy proximally and distally, glabrous, on caudex from fleshy rhizome. |
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Leaves | basal and cauline; basal: 1–6; stipules adnate to petiole, forming 2 linear, membranous wings, wing margins entire or laciniate, each wing with lanceolate to ± deltate projection, margins entire or laciniate, apex acute to long-acuminate; petiole 1.8–14.5 cm, puberulent to tomentose; blade purplish, purple-tinted, or gray-green abaxially, gray, green, or gray-green adaxially, sometimes shiny adaxially, ovate, orbiculate, oblong, deltate, or lanceolate, 0.8–5.3 × 0.4–4.1 cm, often fleshy, base cordate, subcordate, truncate, or attenuate, oblique or not, margins usually ± crenate, serrate, dentate, or coarsely or irregularly repand-dentate, sometimes entire, usually ciliate, apex acute to obtuse or rounded, surfaces glabrous, puberulent, or tomentose; cauline similar to basal except: stipules linear, lanceolate, oblanceolate, or ± oblong to ovate, margins entire, lacerate, or laciniate, usually ciliate, apex sometimes divided into 2–3 filiform processes or obtuse; petiole 0.3–19.7 cm, glabrous or puberulent; blade ovate, oblong, elliptic, deltate, or lanceolate, 0.9–5.2 × 0.2–2.9 cm, length 0.8–7.1 times width, margins crenate, serrate, dentate, repand-denticulate, undulate-denticulate, sinuate, undulate, or entire, abaxial surface puberulent, canescent, or tomentose, adaxial surface glabrous, sparsely pubescent, puberulent, canescent, or tomentose. |
cauline; stipules lanceolate to ovate- or oblong-lanceolate or linear, margins sparingly glandular-fimbriate, apex acute; petiole 2–6 cm, glabrous; blade ovate to ovate-deltate or ovate-lanceolate, 1.2–2.4 × 0.7–1.3 cm, base broadly cuneate to rounded or truncate, margins entire or with 1–3 crenations on proximal 1/2, eciliate, apex acute to rounded, surfaces glabrous, sometimes with a few short hairs on veins abaxially. |
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Peduncles | 3.5–6 cm, glabrous. |
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Flowers | sepals linear to linear-lanceolate, margins eciliate, auricles 0.5–1.5 mm; petals deep lemon-yellow adaxially, upper 2 reddish brown abaxially, lateral 2 and lowest dark brown-veined basally, lateral 2 bearded, lowest 7–11 mm, spur yellow, gibbous, 1–1.4 mm; style head bearded; cleistogamous flowers absent. |
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Capsules | ovoid to ± spherical, 4–7 mm, puberulent. |
ovoid, 3–4.5 mm, glabrous. |
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Seeds | light to dark brown or mottled gray and brown, 2–3.1 mm. |
light brown, ± 2 mm. |
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2n | = 12. |
= 24. |
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Viola purpurea |
Viola guadalupensis |
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Phenology | Flowering May. | |||||||||||||||||||||||||
Habitat | Openings and narrow ledges on limestone rock faces | |||||||||||||||||||||||||
Elevation | 2600 m (8500 ft) | |||||||||||||||||||||||||
Distribution |
AZ; CA; CO; ID; MT; NV; OR; UT; WA; WY; BC; nw Mexico
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TX |
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Discussion | Varieties 7 (7 in the flora). Varieties of Viola purpurea are variable and intergrade. All are found in California; six occur in other western states, one in Mexico, and one in British Columbia. Mature plants are needed for determination. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Viola guadalupensis is known only from the eastern rim of the Guadalupe Mountains in Culberson County. Powell and Wauer noted that it is the only yellow-flowered violet known in the Guadalupe Mountains and appears to be related to V. nuttallii and V. vallicola. K. W. Allred (2008) stated that a report of this species in New Mexico by J. T. Kartesz and C. A. Meacham (1999) requires verification. K. Haskins (pers. comm.) reported that experiments are currently being conducted to propagate plants of V. guadalupensis via cell tissue culture. Chloroplast (trnL-F spacer) and low-copy nuclear gene (GPI) phylogenies indicate that Viola guadalupensis is an alloploid that originated through hybridization between an unidentified member of subsect. Canadenses (the paternal parent) and a member of the V. nuttallii complex (the maternal parent), of sect. Chamaemelanium (T. Marcussen et al. 2011). Evidence reported by these authors from a fossil-calibrated relaxed clock dating analysis showed the estimated maximum age of V. guadalupensis to be (5.7–)8.6(–11.6) million years. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 150. | FNA vol. 6, p. 132. | ||||||||||||||||||||||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||||||||||||||||||||||
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Name authority | Kellogg: Pacific (San Francisco), 2 Feb. 1855: unnumb. (1855) | A. M. Powell & Wauer: Sida 14: 1, fig. 1. (1990) | ||||||||||||||||||||||||
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