Viola pinetorum |
Viola missouriensis |
|||||
---|---|---|---|---|---|---|
goosefoot yellow violet, mountain yellow violet, pine violet |
Missouri violet |
|||||
Habit | Plants perennial, caulescent, not stoloniferous, 3–18 cm, cespitose or not. | Plants perennial, acaulescent, not stoloniferous, 5–50 cm; rhizomes thick, fleshy. | ||||
Stems | 1–3, prostrate or erect, leafy proximally and distally, puberulent or canescent to gray-tomentose, sometimes glabrous, on caudex from subligneous rhizome. |
|||||
Leaves | basal and cauline; basal: 1–4; stipules adnate to petiole forming 2 linear-lanceolate wings, margins entire or laciniate, apex of each wing free, tips usually filamentous; petiole 2.3–9.5 cm, puberulent or canescent; blade purple-tinted abaxially or not, usually linear to narrowly lanceolate, oblanceolate or obovate, or lanceolate-elliptic, rarely ovate, 1.3–5 × 0.3–2.5 cm, base attenuate, margins usually lacerate, dentate, or serrate, sometimes entire, usually undulate, ciliate, apex acute, mucronulate, surfaces puberulent to canescent or gray-tomentose; cauline similar to basal except: stipules lanceolate, oblanceolate, or linear-oblong, margins entire or lacerate, apex acute to acuminate; petiole 0.9–8.3 cm; blade 2.8–9.6 × 0.3–1.4 cm, length 4–11 times width. |
basal, 1–8, ascending to erect; stipules linear-lanceolate to broadly lanceolate, margins entire, sometimes distally glandular, apex acute; petiole 5–20 cm, glabrous; blade green abaxially, unlobed, usually narrowly to broadly deltate, 1.5–12 × 1.5–10 cm, not fleshy, base cordate or broadly cordate to truncate, margins ± crenate to serrate, ciliate or eciliate, apex acute to acuminate, surfaces glabrous, rarely pubescent. |
||||
Peduncles | 2.9–11.5 cm, puberulent or canescent. |
3–25 cm, usually glabrous. |
||||
Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 red- to purple-brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 5–12 mm, spur color same as petals, gibbous, 1.5–3 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to dark blue-violet, lowest and sometimes lateral 2 purple-veined, lateral 2 bearded, lowest beardless, rarely lightly bearded, 15–25 mm, spur same color as petals, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
||||
Capsules | ovoid, 3.5–7 mm, puberulent. |
ellipsoid, 5–12 mm, glabrous. |
||||
Seeds | medium to dark brown, 2–3.5 mm. |
beige, mottled to bronze, 1.5–2.5 mm. |
||||
2n | = 12. |
= 54. |
||||
Viola pinetorum |
Viola missouriensis |
|||||
Phenology | Flowering Mar–May. | |||||
Habitat | Swamps, thickets, stream banks, alluvial woods | |||||
Elevation | 50–2000 m (200–6600 ft) | |||||
Distribution |
CA
|
AL; AR; DC; DE; FL; GA; IA; IL; IN; KS; LA; MD; MN; MO; MS; NC; NE; NJ; NM; OK; SC; SD; TN; TX; VA; WI
|
||||
Discussion | Flowers of Viola pinetorum have been observed to close up in late afternoon then fully reopen the following morning. Although E. O. Wooton and P. C. Standley (1915) reported Viola pinetorum from New Mexico, the plant was probably V. nuttallii. K. W. Allred (2008) noted that V. pinetorum occurs in California; he did not recognize it in New Mexico. Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
What many have treated as Viola affinis, especially in the southern Gulf coastal states, is likely to be V. missouriensis. N. H. Russell (1965) considered the two as likely part of a species complex inhabiting alluvial woods and wet areas and exhibiting the typical deltate leaf blade shape. L. E. McKinney (1992) considered V. missouriensis a variety of V. sororia. Viola missouriensis appears to have a closer affinity to V. affinis, as Russell suggested; current evidence suggests maintaining V. missouriensis as a separate species. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
||||
Key |
|
|||||
Source | FNA vol. 6, p. 144. | FNA vol. 6, p. 138. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. purpurea var. pinetorum | V. candidula, V. langloisii, V. lucidifolia, V. sororia var. missouriensis | ||||
Name authority | Greene: Pittonia 2: 14. (1889) | Greene: Pittonia 4: 141. (1900) | ||||
Web links |