Viola pinetorum |
Viola hirsutula |
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goosefoot yellow violet, mountain yellow violet, pine violet |
southern woodland violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–18 cm, cespitose or not. | Plants perennial, acaulescent, not stoloniferous, 2–15 cm; rhizome thick, fleshy. | ||||
Stems | 1–3, prostrate or erect, leafy proximally and distally, puberulent or canescent to gray-tomentose, sometimes glabrous, on caudex from subligneous rhizome. |
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Leaves | basal and cauline; basal: 1–4; stipules adnate to petiole forming 2 linear-lanceolate wings, margins entire or laciniate, apex of each wing free, tips usually filamentous; petiole 2.3–9.5 cm, puberulent or canescent; blade purple-tinted abaxially or not, usually linear to narrowly lanceolate, oblanceolate or obovate, or lanceolate-elliptic, rarely ovate, 1.3–5 × 0.3–2.5 cm, base attenuate, margins usually lacerate, dentate, or serrate, sometimes entire, usually undulate, ciliate, apex acute, mucronulate, surfaces puberulent to canescent or gray-tomentose; cauline similar to basal except: stipules lanceolate, oblanceolate, or linear-oblong, margins entire or lacerate, apex acute to acuminate; petiole 0.9–8.3 cm; blade 2.8–9.6 × 0.3–1.4 cm, length 4–11 times width. |
basal, 2–4, prostrate or nearly so; stipules linear-lanceolate, margins entire, apex acute; petiole 1–10 cm, usually glabrous; blade unlobed, reniform to ovate, 1–6 × 1–5 cm, base cordate, margins serrate, ciliate or eciliate, apex rounded, adaxial leaf surface with silvery strigose patches. |
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Peduncles | 2.9–11.5 cm, puberulent or canescent. |
2–12 cm, glabrous. |
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Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 red- to purple-brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 5–12 mm, spur color same as petals, gibbous, 1.5–3 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm; petals light to deep reddish violet to blue-violet on both surfaces, 3 lower whitish basally, dark violet-veined, lateral 2 densely bearded, lowest 11–17 mm, spur violet, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
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Capsules | ovoid, 3.5–7 mm, puberulent. |
ellipsoid, 8–12 mm, glabrous. |
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Seeds | medium to dark brown, 2–3.5 mm. |
beige, mottled to bronze, 1–1.5 mm. |
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2n | = 12. |
= 54. |
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Viola pinetorum |
Viola hirsutula |
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Phenology | Flowering Mar–Jun. | |||||
Habitat | Dry to mesic deciduous woodlands | |||||
Elevation | 100–1000 m (300–3300 ft) | |||||
Distribution |
CA
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AL; CT; DC; DE; FL; GA; IN; KY; MD; MS; NC; NJ; NY; OH; PA; SC; TN; VA; WV
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Discussion | Flowers of Viola pinetorum have been observed to close up in late afternoon then fully reopen the following morning. Although E. O. Wooton and P. C. Standley (1915) reported Viola pinetorum from New Mexico, the plant was probably V. nuttallii. K. W. Allred (2008) noted that V. pinetorum occurs in California; he did not recognize it in New Mexico. Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Viola hirsutula exhibits little phenotypic plasticity. Although considered by N. L. Gil-Ad (1997) to be a putative hybrid, he did not identify the potential parents and chose not to recognize it. A. Cronquist in H. A. Gleason and Cronquist (1991) considered V. hirsutula a form of V. villosa. Viola hirsutula is considered one of the more distinct species throughout its range by McKinney because of the pattern of silvery strigose patches on adaxial leaf surfaces and a more prostrate growth habit than similar species. He believes V. hirsutula is one of the least variable species in the flora. Viola hirsutula appears to occur much less frequently than herbarium collections suggest. Some herbarium specimens are misidentified and represent depauperate plants, usually of V. sororia. These specimens are often hairy on the adaxial leaf surfaces but lack the silvery patches. Viola hirsutula reportedly hybridizes with V. sororia var. sororia (= V. ×cordifolia [Nuttall] Schweinitz) and V. sagittata var. sagittata (= V. ×redacta House). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 144. | FNA vol. 6, p. 133. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. purpurea var. pinetorum | |||||
Name authority | Greene: Pittonia 2: 14. (1889) | Brainerd: Rhodora 9: 98. (1907) | ||||
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