Viola pinetorum
Viola guadalupensis
goosefoot yellow violet, mountain yellow violet, pine violet
Guadalupe Mountains violet, Guadalupe violet
1–3, prostrate or erect, leafy proximally and distally, puberulent or canescent to gray-tomentose, sometimes glabrous, on caudex from subligneous rhizome.
1–5, decumbent to erect, leafy proximally and distally, glabrous, on caudex from fleshy rhizome.
basal and cauline;
basal: 1–4;
stipules adnate to petiole forming 2 linear-lanceolate wings, margins entire or laciniate, apex of each wing free, tips usually filamentous;
petiole 2.3–9.5 cm, puberulent or canescent;
blade purple-tinted abaxially or not, usually linear to narrowly lanceolate, oblanceolate or obovate, or lanceolate-elliptic, rarely ovate, 1.3–5 × 0.3–2.5 cm, base attenuate, margins usually lacerate, dentate, or serrate, sometimes entire, usually undulate, ciliate, apex acute, mucronulate, surfaces puberulent to canescent or gray-tomentose;
cauline similar to basal except: stipules lanceolate, oblanceolate, or linear-oblong, margins entire or lacerate, apex acute to acuminate;
petiole 0.9–8.3 cm;
blade 2.8–9.6 × 0.3–1.4 cm, length 4–11 times width.
cauline;
stipules lanceolate to ovate- or oblong-lanceolate or linear, margins sparingly glandular-fimbriate, apex acute;
petiole 2–6 cm, glabrous;
blade ovate to ovate-deltate or ovate-lanceolate, 1.2–2.4 × 0.7–1.3 cm, base broadly cuneate to rounded or truncate, margins entire or with 1–3 crenations on proximal 1/2, eciliate, apex acute to rounded, surfaces glabrous, sometimes with a few short hairs on veins abaxially.
2.9–11.5 cm, puberulent or canescent.
3.5–6 cm, glabrous.
sepals lanceolate, margins ciliate, auricles 0.5–1 mm;
petals deep lemon-yellow adaxially, upper 2 red- to purple-brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 5–12 mm, spur color same as petals, gibbous, 1.5–3 mm;
style head bearded; cleistogamous flowers axillary.
sepals linear to linear-lanceolate, margins eciliate, auricles 0.5–1.5 mm;
petals deep lemon-yellow adaxially, upper 2 reddish brown abaxially, lateral 2 and lowest dark brown-veined basally, lateral 2 bearded, lowest 7–11 mm, spur yellow, gibbous, 1–1.4 mm;
style head bearded; cleistogamous flowers absent.
ovoid, 3.5–7 mm, puberulent.
ovoid, 3–4.5 mm, glabrous.
medium to dark brown, 2–3.5 mm.
light brown, ± 2 mm.
= 12.
= 24.
Viola pinetorum
Viola guadalupensis
Flowers of Viola pinetorum have been observed to close up in late afternoon then fully reopen the following morning.
Although E. O. Wooton and P. C. Standley (1915) reported Viola pinetorum from New Mexico, the plant was probably V. nuttallii. K. W. Allred (2008) noted that V. pinetorum occurs in California; he did not recognize it in New Mexico.
Varieties 2 (2 in the flora).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Of conservation concern.
Viola guadalupensis is known only from the eastern rim of the Guadalupe Mountains in Culberson County. Powell and Wauer noted that it is the only yellow-flowered violet known in the Guadalupe Mountains and appears to be related to V. nuttallii and V. vallicola. K. W. Allred (2008) stated that a report of this species in New Mexico by J. T. Kartesz and C. A. Meacham (1999) requires verification. K. Haskins (pers. comm.) reported that experiments are currently being conducted to propagate plants of V. guadalupensis via cell tissue culture.
Chloroplast (trnL-F spacer) and low-copy nuclear gene (GPI) phylogenies indicate that Viola guadalupensis is an alloploid that originated through hybridization between an unidentified member of subsect. Canadenses (the paternal parent) and a member of the V. nuttallii complex
(the maternal parent), of sect. Chamaemelanium (T. Marcussen et al. 2011). Evidence reported by these authors from a fossil-calibrated relaxed clock dating analysis showed the estimated maximum age of V. guadalupensis to be (5.7–)8.6(–11.6) million years.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
1. Plants 6.5–18 cm, not cespitose; basal leaf blades 0.7–2.5 cm wide, surfaces puberulent; peduncles 3.4–11.5 cm. | var. pinetorum |
1. Plants 3–7(–9) cm, usually cespitose; basal leaf blades 0.3–1 cm wide, surfaces canescent, sometimes appearing gray-tomentose; peduncles 2.9–6(–7) cm. | var. grisea |
CA
WildflowerSearch
iNaturalist
USDA Plants Database
