Viola pinetorum |
Viola clauseniana |
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goosefoot yellow violet, mountain yellow violet, pine violet |
Clausen's violet |
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Habit | Plants perennial, caulescent, not stoloniferous, 3–18 cm, cespitose or not. | Plants perennial, acaulescent, not stoloniferous, 3–20 cm; rhizome short, slender, fleshy. | ||||
Stems | 1–3, prostrate or erect, leafy proximally and distally, puberulent or canescent to gray-tomentose, sometimes glabrous, on caudex from subligneous rhizome. |
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Leaves | basal and cauline; basal: 1–4; stipules adnate to petiole forming 2 linear-lanceolate wings, margins entire or laciniate, apex of each wing free, tips usually filamentous; petiole 2.3–9.5 cm, puberulent or canescent; blade purple-tinted abaxially or not, usually linear to narrowly lanceolate, oblanceolate or obovate, or lanceolate-elliptic, rarely ovate, 1.3–5 × 0.3–2.5 cm, base attenuate, margins usually lacerate, dentate, or serrate, sometimes entire, usually undulate, ciliate, apex acute, mucronulate, surfaces puberulent to canescent or gray-tomentose; cauline similar to basal except: stipules lanceolate, oblanceolate, or linear-oblong, margins entire or lacerate, apex acute to acuminate; petiole 0.9–8.3 cm; blade 2.8–9.6 × 0.3–1.4 cm, length 4–11 times width. |
basal, 2–10, often prostrate, sometimes ascending; stipules narrowly lanceolate, margins faintly glandular-toothed, apex acute or obtuse; petiole 5–11 cm, glabrous; blade unlobed, ± deltate, 3–5 × 4–5 cm, base ± truncate, margins serrate, ciliate or eciliate, apex obtuse, surfaces usually glabrous, sometimes sparsely pubescent abaxially. |
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Peduncles | 2.9–11.5 cm, puberulent or canescent. |
8–14 cm, glabrous. |
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Flowers | sepals lanceolate, margins ciliate, auricles 0.5–1 mm; petals deep lemon-yellow adaxially, upper 2 red- to purple-brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 5–12 mm, spur color same as petals, gibbous, 1.5–3 mm; style head bearded; cleistogamous flowers axillary. |
sepals lanceolate to ovate, margins eciliate or ciliate around auricles, auricles 1–2 mm; petals light violet on both surfaces, lower 3 purple-veined, all beardless, lowest petal 8–20 mm, spur white, gibbous, 2–3 mm; style head beardless; cleistogamous flowers on prostrate to ascending peduncles. |
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Capsules | ovoid, 3.5–7 mm, puberulent. |
oblong, 8–10 mm, glabrous. |
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Seeds | medium to dark brown, 2–3.5 mm. |
dark brown to black, 1.2–2 mm. |
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2n | = 12. |
= 44. |
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Viola pinetorum |
Viola clauseniana |
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Phenology | Flowering Apr–Jun. | |||||
Habitat | Hanging gardens, seeps, springs, shady areas | |||||
Elevation | 1600 m (5200 ft) | |||||
Distribution |
CA
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UT |
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Discussion | Flowers of Viola pinetorum have been observed to close up in late afternoon then fully reopen the following morning. Although E. O. Wooton and P. C. Standley (1915) reported Viola pinetorum from New Mexico, the plant was probably V. nuttallii. K. W. Allred (2008) noted that V. pinetorum occurs in California; he did not recognize it in New Mexico. Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Of conservation concern. Viola clauseniana is endemic to Zion National Park, Washington County. M. S. Baker reported that its seeds were minutely roughed, a characteristic not recorded for other Viola species. Viola clauseniana was originally thought to be closely related to the acaulescent blue violets, most notably V. nephrophylla (S. L. Welsh et al. 1987; L. E. McKinney 1992). After contemplating the 2n = 44 chromosome count obtained by J. Clausen (1964), H. E. Ballard (pers. comm.) suggested that V. clauseniana might be more closely related with the stemless white violets (for example, V. blanda) than with V. nephrophylla. Viola clauseniana was considered a distinct species by N. H. Holmgren (2005d) and T. Marcussen and T. Karlsson (2010). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 6, p. 144. | FNA vol. 6, p. 128. | ||||
Parent taxa | Violaceae > Viola | Violaceae > Viola | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | V. purpurea var. pinetorum | |||||
Name authority | Greene: Pittonia 2: 14. (1889) | M. S. Baker: Madroño 4: 194. (1938) | ||||
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