Viola pedunculata
Viola tripartita
California golden violet, johnny-jump-up, wild pansy, yellow pansy
three-part or Piedmont or threepart violet, threepart violet
1–10+, decumbent, ascending, or erect, leafy proximally and distally, glabrous or puberulent, from shallow to deep-seated, enlarged rhizome with fleshy to subligneous roots.
1(2), erect, leafless proximally, leafy distally, glabrous or puberulent, from subligneous rhizome.
cauline;
stipules ovate, linear-lanceolate, or oblanceolate, sometimes leaflike, margins entire or glandular-toothed, apex acute to acuminate;
petiole 2.7–7.2 cm, usually finely puberulent, sometimes glabrate;
blade deltate to ovate, 1–5.5 × 1–5.5 cm, base truncate, subcordate, or attenuate, margins crenate to serrate, ciliate, apex acute to obtuse, surfaces subglabrous or sparsely puberulent.
basal and cauline;
basal: 0(–2);
stipules ovate, not leaflike, margins entire, apex acute to acuminate, surfaces glabrous or puberulent;
petiole 9–11.5 cm, glabrous or puberulent;
blade unlobed, ovate, or 3–5-lobed, 4–5 × 1–5 cm, base cordate, margins entire or crenate-serrate, ciliate or eciliate, apex acute, surfaces glabrous or ± puberulent;
cauline similar to basal except: restricted to distal ends of stems;
stipules ovate to oblong;
petiole 0.7–7.2 cm, glabrous or puberulent;
blade unlobed, ovate or deltate, or 3-lobed (if 3-lobed, lateral lobes falcate, middle rhombic, longer than others, lobes may appear petiolate; unlobed and 3-lobed leaves can occur on same plant), 1–6 × 0.5–5.5 cm, base truncate to cuneate, margins serrate, ciliate or eciliate, surfaces glabrous or pubescent.
2.9–20 cm, sparsely to densely puberulent.
1.5–4 cm, glabrous or pubescent.
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–3 mm;
petals golden yellow adaxially, upper 2 reddish brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 10–20 mm, spur dark reddish brown, gibbous, 2–4 mm;
style head bearded; cleistogamous flowers absent.
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 0.1–0.5 mm;
petals lemon-yellow adaxially, upper 2, rarely others, brownish purple abaxially, lowest and usually lateral 2 brownish purple-veined, lateral 2 and sometimes lowest bearded, lowest 10–18 mm, spur yellow, gibbous, 0.5–2 mm;
style head bearded; cleistogamous flowers axillary.
ellipsoid, 5–11 mm, glabrous.
ovoid to ellipsoid, 9–12 mm, glabrous.
dark brown or black, shiny, 2.7 mm.
beige, bronze, or brown, 2.4–3 mm.
= 12.
= 12.
Viola pedunculata
Viola tripartita
The stems of Viola pedunculata arise from an enlarged, subterranean, spongy or fibrous rhizome. Often, these rhizome structures are deep seated; it is unknown how they get so deeply buried. The anther appendages of V. pedunculata are hairy distally, a characteristic not known to occur in other members of the V. purpurea complex.
Larvae of the federally listed Callippe silverspot butterfly [Speyeria callippe (Boisduval) callippe] feed only on Viola pedunculata.
Plants with leaves reported to be smaller, thinner, deltate, mostly longer than wide, with yellow petals (versus orange for Viola pedunculata var. pedunculata), style 2.1 mm (versus 2.9 mm for var. pedunculata), from the Pinnacles region in San Benito County, California, have been called subsp. tenuifolia.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
Some authors recognize two varieties of Viola tripartita based on lobed versus unlobed leaves. F. L. Lévesque and P. M. Dansereau (1966) suggested that leaf variation is the only character difference between vars. tripartita and glaberrima. N. H. Russell (1965) stated that V. tripartita plants with lobed and unlobed leaves are sympatric and frequently intergrade and did not recognize them as distinct. The situation with two leaf forms in V. tripartita is similar to V. lobata, which also has two leaf forms.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
CA
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