Viola pedunculata
Viola subsinuata
California golden violet, johnny-jump-up, wild pansy, yellow pansy
early blue violet, wavy-leaf violet, wood violet
1–10+, decumbent, ascending, or erect, leafy proximally and distally, glabrous or puberulent, from shallow to deep-seated, enlarged rhizome with fleshy to subligneous roots.
cauline;
stipules ovate, linear-lanceolate, or oblanceolate, sometimes leaflike, margins entire or glandular-toothed, apex acute to acuminate;
petiole 2.7–7.2 cm, usually finely puberulent, sometimes glabrate;
blade deltate to ovate, 1–5.5 × 1–5.5 cm, base truncate, subcordate, or attenuate, margins crenate to serrate, ciliate, apex acute to obtuse, surfaces subglabrous or sparsely puberulent.
basal, 2–11, ascending to erect;
stipules linear-lanceolate, margins entire, apex acute;
petiole 5–25 cm, glabrous or pubescent;
mid-season leaf blades incised or lobed throughout, earliest leaf blades lobed (plants homophyllous), similar to mid-season blades, blade 5–9(–16)-lobed, sinuses usually narrower, shallower toward leaf base, middle and lateral blade lobes differ in width and/or shape, middle lobes narrowly deltate to narrowly elliptic, lateral lobes narrowly elliptic, lanceolate, or falcate, 1–11 × 1–12 cm, base truncate to cordate, margins entire or crenate, ciliate or eciliate, apex acute, mucronulate, surfaces usually pubescent throughout or on veins.
2.9–20 cm, sparsely to densely puberulent.
5–15 cm, glabrous or pubescent.
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–3 mm;
petals golden yellow adaxially, upper 2 reddish brown abaxially, lower 3 dark brown-veined, lateral 2 bearded, lowest 10–20 mm, spur dark reddish brown, gibbous, 2–4 mm;
style head bearded; cleistogamous flowers absent.
sepals lanceolate to ovate, margins ciliate or eciliate, auricles 1–2 mm;
petals light to dark blue-violet on both surfaces, lower 3 and upper 2 sometimes purple-veined, lateral 2 bearded, lowest sometimes bearded, 15–25 mm, spur color same as petals, gibbous, 2–3 mm;
style head beardless; cleistogamous flowers on prostrate to ascending peduncles.
ellipsoid, 5–11 mm, glabrous.
ellipsoid, 8–12 mm, glabrous.
dark brown or black, shiny, 2.7 mm.
beige, mottled to bronze, 1.5–2.5 mm.
= 12.
= 54.
Viola pedunculata
Viola subsinuata
The stems of Viola pedunculata arise from an enlarged, subterranean, spongy or fibrous rhizome. Often, these rhizome structures are deep seated; it is unknown how they get so deeply buried. The anther appendages of V. pedunculata are hairy distally, a characteristic not known to occur in other members of the V. purpurea complex.
Larvae of the federally listed Callippe silverspot butterfly [Speyeria callippe (Boisduval) callippe] feed only on Viola pedunculata.
Plants with leaves reported to be smaller, thinner, deltate, mostly longer than wide, with yellow petals (versus orange for Viola pedunculata var. pedunculata), style 2.1 mm (versus 2.9 mm for var. pedunculata), from the Pinnacles region in San Benito County, California, have been called subsp. tenuifolia.
(Discussion copyrighted by Flora of North America; reprinted with permission.)
L. E. McKinney (1992) described the misconceptions surrounding Viola subsinuata that many came to think of as the heterophyllous V. palmata, whereas V. subsinuata is homophyllous. As E. Brainerd (1910, 1921) pointed out, heterophylly versus homophylly is an important and steadfast character difference in acaulescent Viola species. N. L. Gil-Ad (1997) chose not to recognize V. subsinuata, suggesting that it represented a hybrid or introgressant. H. E. Ballard (2000) and A. Haines (2011) both recognized V. subsinuata as a species.
Viola subsinuata is a state historical species in Rhode Island, where it was last documented in 1941 (R. W. Enser, http://rinhs.org/wp-content/uploads/ri_rare_plants_2007.pdf).
(Discussion copyrighted by Flora of North America; reprinted with permission.)
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